Oxymeris modesta ( Defrance, 1829 )

Harzhauser, Mathias & Landau, Bernard M., 2023, The auger snails (Gastropoda, Conoidea, Terebridae) of the Miocene Paratethys Sea, Zootaxa 5385 (1), pp. 1-70 : 26-30

publication ID

https://doi.org/ 10.11646/zootaxa.5385.1.1

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lsid:zoobank.org:pub:7FBEC9FB-E31E-4CA4-8BD0-BE6D35322C3E

DOI

https://doi.org/10.5281/zenodo.10463088

persistent identifier

https://treatment.plazi.org/id/7C37CE5F-FFB9-AB28-FF2C-FCEC32DDFA8E

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scientific name

Oxymeris modesta ( Defrance, 1829 )
status

 

Oxymeris modesta ( Defrance, 1829)

Figs 13A–F View FIGURE 13

* Terebra modesta View in CoL Tristan — Defrance 1829: 288.

Subula Blainvillii View in CoL m.— Eichwald 1830: 223.

Terebra Blainvillei [sic] m— Eichwald 1851: 89, pl. 7, figs 23a–b.

Terebra Blainvillei [sic]— Eichwald 1852: plate captions, pl. 7, figs 23a–b.

Terebra Blainvillei [sic] m.— Eichwald 1853: 161.

Terebra fuscata Brn. — Hauer 1837: 417 [non Oxymeris fuscata ( Brocchi, 1814) ].

Terebra fuscata — Hörnes 1848: 17 [non Oxymeris fuscata ( Brocchi, 1814) ].

Terebra fuscata —Naumann 1852: pl. 79, fig. 22 [non Oxymeris fuscata ( Brocchi, 1814) ].

Terebra fuscata Brocc. — Hörnes 1852: 128, pl. 11, figs 15–18, 26 [non Oxymeris fuscata ( Brocchi, 1814) ].

Terebra fuscata Brocc. — Pereira da Costa 1866: 78, pl. 12, figs 14–16, pl. 13, figs 1–2 [non Oxymeris fuscata ( Brocchi, 1814) ].

Terebra Hochstetteri View in CoL —Hoernes 1880: 246 [nomen nudum].

Terebra (Acus) Hochstetteri nov. form.— Hoernes & Auinger 1880: 107, pl. 12, figs 18 a–b [non Terebra hochstetteri Martin, 1879 View in CoL ].

Terebra Hochstetteri View in CoL ??— Hoernes & Auinger 1880: 107, pl. 12, fig. 19.

[ Terebra (Subula) plicaria View in CoL ] var. perstriatula Sacc. — Sacco 1891a: 14 [pro Terebra hochstetteri Hoernes & Auinger 1880 View in CoL : pl. 12, fig. 19].

[Terebra] S [ubula]. plicaria var. laevisulcata Sacc. — Sacco 1891a: 12, pl. 1, fig. 14.

[Terebra] Subula conicoplicaria Sacc. — Sacco 1891a: 15, pl. 1, fig. 23.

[Terebra] Subula modesta ( Tristan )— Sacco 1891a: 15, pl. 1, fig. 24.

[ Terebra] S [ubula]. modesta var. plano-clavata Sacc.— Sacco 1891a: 16, pl. 1, fig. 25.

[Terebra] S [ubula]. modesta var. perinflata Sacc. — Sacco 1891a: 16, pl. 1, fig. 26.

[Terebra] S [ubula]. modesta var. dertofusulata Sacc. — Sacco 1891a: 16, pl. 1, fig. 27.

[Terebra] S [ubula]. modesta var. ovulata Sacc. — Sacco 1891a: 16, pl. 1, fig. 28.

Terebra (Acus) fuscata Brocc. — Boettger 1902: 18 [non Oxymeris fuscata ( Brocchi, 1814) ].

Terebra (Subula) fuscata Brocc. — Friedberg 1911: 1, pl. 1, fig. 1 [non Oxymeris fuscata ( Brocchi, 1814) ].

Terebra (Subula) modesta Tristan in Defrance View in CoL — Peyrot 1931: 121, pl. 10, fig. 20.

Terebra modesta Trist. View in CoL — Friedberg 1938: 158.

Terebra (Subula) plicaria Bast. View in CoL — Csepreghy-Meznerics 1950: 62 [non Oxymeris plicaria (de Basterot, 1825) View in CoL ].

Terebra (Oxymeris) modesta View in CoL Tristan in Defrance, 1829 — Glibert 1952a: 380, pl. 14, fig. 5.

Terebra (Subula) plicaria Bast. View in CoL — Csepreghy-Meznerics 1954: 57 [non Oxymeris plicaria (de Basterot, 1825) View in CoL ].

Terebra (Subula) fuscata plicaria Bast. View in CoL — Strausz 1954: 37, pl. 2, figs 44a–b [non Oxymeris plicaria (de Basterot, 1825) View in CoL ].

Terebra (Subula) fuscata Brocc. — Korobkov 1955: plate captions, pl. 92, figs 7a, 7b.

Terebra (Subula) plicaria Bast. View in CoL — Csepreghy-Meznerics 1956: pl. 12, figs 5–6 [non Oxymeris plicaria (de Basterot, 1825) View in CoL ].

S [ubula]. (O [xymeris].) fuscata plicaria (Bast.) View in CoL — Sieber 1958: 162 [non Oxymeris plicaria (de Basterot, 1825) View in CoL ].

S [ubula]. (O [xymeris].) fuscata modesta (Trist.) — Sieber 1958: 162.

Terebra (Subula) fuscata (Brocchi) — Erünal-Erentöz 1958: 125, pl. 20, figs 15, 16 [non Oxymeris fuscata ( Brocchi, 1814) ].

Terebra (Subula) fuscata var. modesta View in CoL Tristan in Defrance, 1829 —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 218, pl. 52, figs 1–2.

Terebra (Subula) fuscata plicaria Basterot View in CoL — Strausz 1962: 31, pl. 5, figs 8–10 [non Oxymeris plicaria (de Basterot, 1825) View in CoL ].

Terebra (Subula) fuscata Brocchi, 1814 — Strausz 1966: 394, pl. 5, fig. 11.

Terebra (Subula) fuscata plicaria Basterot View in CoL — Strausz 1966: 394, pl. 5, figs 8–10 [non Oxymeris plicaria (de Basterot, 1825) View in CoL ].

Subula (Oxymeris) fuscata (Brocchi, 1914) — Tejkal et al. 1967: 209, pl. 12B, fig. 3 [non Oxymeris fuscata ( Brocchi, 1814) ].

Terebra fuscata ( Brocchi, 1814) — Zelinskaya et al. 1968: 231, pl. 52, figs 6–7 [non Oxymeris fuscata ( Brocchi, 1814) ].

Terebra (Subula) fuscata Brocchi — Atanacković 1969: 216, pl. 14, figs 2–2a [non Oxymeris fuscata ( Brocchi, 1814) ].

Subula (Oxymeris) fuscata modesta (Tristan, 1829)— Steininger et al. 1971: 406, pl. 11, fig. 11.

Terebra (Subula) plicaria Basterot View in CoL — Csepreghy-Meznerics 1972: 34, pl. 18, figs 4–5 [non Oxymeris plicaria (de Basterot, 1825 )].

Terebra (Subula) fuscata Brocchi — Nicorici 1972: 69 [non Oxymeris fuscata ( Brocchi, 1814) ].

Terebra (Oxymeris) modesta (Tristan in Defrance)— Bohn-Havas 1973: 1126, pl. 8, fig. 7.

Terebra (Subula) plicaria (Basterot) 1825 — Bohn-Havas 1973: 1126 [non Oxymeris plicaria (de Basterot, 1825) ].

Subula (Subula) plicaria (Basterot) — Davoli 1977: 156, pl. 3, figs 1–5, 9, 19.

Subula (Oxymeris) modesta (Tristan in Defrance)— Davoli, 1977: 158, pl. 3, figs 6–8, 10, 13, 17, 25.

Terebra (Subula) fuscata plicaria Basterot, 1825 — Švagrovský 1982: 408, pl. 7, figs 1–3 [non Oxymeris plicaria (de Basterot, 1825) ].

Subula (Subula) modesta Tristan in Defrance— Ruggieri & Davoli 1984: 73, pl. 15, figs 4–6.

Terebra (Subula) fuscata modesta Tristan in Defrance, 1817— Atanacković 1985: 183, pl. 41, figs 5–6.

Subula (Oxymeris) plicaria ( Basterot, 1825) — Bałuk 1997: 68, pl. 24, figs 6–12 [non Oxymeris plicaria (de Basterot, 1825) ].

Subula (Subula) fuscata plicaria (Basterot) — Schultz 1998: 72, pl. 29, fig. 14 [non Oxymeris plicaria (de Basterot, 1825) ].

Subula (Oxymeris) plicaria ( Basterot, 1825) —Harzhauser 2002: 115, pl. 10, fig. 7.

Subula (Oxymeris) plicaria ( Basterot 1825) — İslamoğlu 2004: 165, pl. 4, figs 11, 12 [non Oxymeris plicaria (de Basterot, 1825)].

Subula plicaria (de Basterot, 1825) — Landau et al. 2013: 296, pl. 51, figs 8–9; pl. 82, fig. 15 [non Oxymeris plicaria (de Basterot, 1825)].

Subula (Subula) fuscata plicaria ( Basterot, 1825) — Popa et al. 2014: 18, pl. 6, fig. 2 [non Oxymeris plicaria (de Basterot, 1825)].

non Terebra (Subula) fuscata modesta Trist. — Strausz 1954: 37, pl. 2, figs 45a–c [= Oxymeris transleithana nov. sp.].

non Terebra (Subula) fuscata modesta Tristan— Strausz 1962: 31, pl. 5, figs 12–14 [= Oxymeris transleithana nov. sp.].

non Terebra (Subula) fuscata modesta Tristan (in Defrance)— Strausz 1966: 395, pl. 5, figs 12–14 [= Oxymeris transleithana nov. sp.].

Type material. The syntypes of Oxymeris modesta were described from the Langhian of the Touraine ( France). The collection of Defrance was destroyed in WWII (pers. comm. Ronald Janssen) .

Illustrated material. NHMW 1855/0045/0396, 105.6 mm, MD: mm, 22.1 mm, Baden ( Austria), Figs 13A View FIGURE 13 1 – A View FIGURE 1 2 View FIGURE 2 . NHMW 2023/0296/0002, SL: 88.1 mm, MD: mm, 23.9 mm, Lăpugiu de Sus ( Romania), Figs 13B View FIGURE 13 1 –B View FIGURE 1 2 View FIGURE 2 . NHMW 1870/0033/0047, 83.1 mm, MD: mm, 22.3 mm, Lăpugiu de Sus ( Romania), Figs 13C View FIGURE 13 1 –C View FIGURE 1 2 View FIGURE 2 .

NHMW 1949/0005/0005, SL: 55.0 mm, MD: 14.8 mm, holotype of Terebra (Acus) hochstetteri Hoernes & Auinger, 1880 ; illustrated in Hoernes & Auinger (1880: pl. 12, fig. 18), Lăpugiu de Sus ( Romania), Figs 13D View FIGURE 13 1 – D View FIGURE 1 2 View FIGURE 2 . NHMW 2023/0296/0003, SL: 88.9 mm, MD: 22.5 mm, Lăpugiu de Sus ( Romania), Figs 13E View FIGURE 13 1 –E View FIGURE 1 2 View FIGURE 2 . NHMW 1843/0008/0010, SL: 119.2 mm, MD: 26.5 mm, Sainte Maure, Touraine ( France), Figs 13F View FIGURE 13 1 –F View FIGURE 1 2 View FIGURE 2 .

Additional material. NHMW 1846 View Materials /0037/0118, SL: 85.2 mm, MD: 22.3 mm, Niederkreuzstetten ( Austria), illustrated in Hörnes (1852; pl. 11, Fig. 15 View FIGURE 15 ); note that the illustration shows a restored specimen with apex . NHMW 1860 View Materials /00050/0047, Neuruppersdorf ( Austria) ; NHMW 1869 View Materials /0001/0026, SL: 74.4 mm, MD: 20.4 mm, Bad Vöslau ( Austria), illustrated in Hörnes (1852; pl. 11, Fig. 18 View FIGURE 18 ); 4 spec., 15 spec. , NHMW 1997 View Materials z0178/1206, Bad Vöslau ( Austria); 1 spec. , NHHMW 1866 /0001/0694, Möllersdorf ( Austria); 20 spec. , NHMW 1997 View Materials z0178/1485, Gainfarn ( Austria); 26 spec. , NHMW 2013 View Materials /0479/1405, Gainfarn ( Austria); 9 spec. , NHMW 1846 View Materials /0037/0117, Bad Vöslau ( Austria); 2 spec. , NHMW 2013 View Materials /0078/0419, Baden ( Austria); 4 spec. , NHMW 2013 View Materials /0300/0402, Baden-Sooss ( Austria); 1 spec. , NHMW 2013 View Materials /0300/0403, Baden-Sooss ( Austria); 3 spec. , NHMW 1853 View Materials /0010/0011, Enzesfeld ( Austria); 8 spec. , NHMW 1869 View Materials /0001/0450, Forchtenau ( Austria); 2 spec. , NHMW 1863 View Materials /0015/0425, Forchtenau ( Austria); 7 spec. , NHMW 1870 View Materials /0037/0009, Ritzing ( Austria); 8 spec. , NHMW 1868 View Materials /0008/0047, Vienna / Pötzleinsdorf ( Austria); 1 spec. , NHMW 2012 View Materials /0229/0169, Traiskirchen ( Austria); 1 spec. , NHMW 2012 View Materials /0229/0170, Traiskirchen ( Austria); 4 spec. , NHMW 1860 View Materials /0001/0136, Steinebrunn ( Austria); 12 spec. , NHMW 1860 View Materials /0005/0028, Pöls ( Austria); 7 spec. , NHMW 1863 View Materials /0015/0927, Hrušovany nad Jevišovkou ( Czech Republic); 12 spec. , NHMW 1851 View Materials /0010/0017, Mikulov ( Czech Republic); 3 spec. , NHMW 1860 View Materials /0001/90137, Mikulov ( Czech Republic); 2 spec. , NHMW 1859 View Materials /0040/0008, Zaleśce ( Ukraine); 1. spec ., NHMW 2016 View Materials /0177/0932, Lăpugiu de Sus ( Romania); 2. spec ., NHMW 2016 View Materials /0177/0928, Lăpugiu de Sus ( Romania); 6. spec ., NHMW 1973 View Materials /1615/0012, Lăpugiu de Sus ( Romania); 8. spec ., NHMW 1870 View Materials /0033/0047, Lăpugiu de Sus ( Romania); 7. spec ., NHMW 1854 View Materials /0035/0310, Lăpugiu de Sus ( Romania); 10. spec ., NHMW 1854 View Materials /0035/0109, Lăpugiu de Sus ( Romania); 10. spec ., NHMW 2016 View Materials /0177/0927, Lăpugiu de Sus ( Romania); 15. spec ., NHMW 2016 View Materials /0177/0926, Lăpugiu de Sus ( Romania); 7. spec ., NHMW 2016 View Materials /0177/0934, Lăpugiu de Sus ( Romania); 8. spec ., NHMW 2016 View Materials /0177/0931, Lăpugiu de Sus ( Romania); 28. spec ., NHMW 2016 View Materials /0177/0930, Lăpugiu de Sus ( Romania); 5 spec. , NHMW 1867 View Materials /0019/0050, CoŞteiu de Sus ( Romania); 2 spec. , NHMW 2018 View Materials /0245/0351, CoŞteiu de Sus ( Romania); 4 spec. , NHMW 2018 View Materials /0245/0352, CoŞteiu de Sus ( Romania) .

Revised description. Very large, moderately slender shell of up to 14 teleoconch whorls; apical angle ~17°. Protoconch unknown. Early spire typically conical or weakly gradate. Whorl profile conical to subcylindrical. Broad subsutural band with weak axial ribs; delimited by distinct spiral groove. Below, broad, low, faintly opisthocyrt axial ribs. Axial sculpture fading abapically, typically around tenth teleoconch whorl, but variable. Spiral grove weakening but frequently persisting as weak groove on last whorl. Later teleoconch whorls weakly convex, usually smooth, except for delicate growth lines. Spiral threads or grooves may occur on last teleoconch whorls. Last whorl high, weakly convex, about 35–40% of total height. Base slowly contracting. Fasciole prominent, broad, adapically delimited by sharp carina and separated from base by moderately narrow groove. Aperture elongate, moderately narrow. Columella weakly twisted, angulated at transition into convex parietal area. Columellar callus forming thin, broad rim, moderately delimited from base. Anal canal indistinct, narrowly incised. Outer lip thin. Basal lip convex, wide. Siphonal canal short, wide, twisted, slightly deflected to the left, moderately notched at tip.

Paratethyan synonyms. Subula blainvillii Eichwald, 1830 , the type is stored in the Paleontological Museum of the St. Petersburg State University ( Russia).

Terebra (Acus) hochstetteri Hoernes & Auinger, 1880 ; holotype, NHMW 1949/0005/0005, SL: 55.0 mm, MD: 14.8 mm, illustrated in Hoernes & Auinger 1880: pl. 12, fig. 18, Lăpugiu de Sus ( Romania), Figs 13D View FIGURE 13 1 –D View FIGURE 1 2 View FIGURE 2 . Terebra (Acus) hochstetteri Hoernes & Auinger, 1880 . Terebra hochstetteri is based on a single specimen with spiral grooves on the last whorl. Although rare, spiral grooves occasionally occur on the last whorls of Oxymeris modesta and do not justify a separation at species level. Terebra hochstetteri is preoccupied by Martin (1879) for a Miocene species from Indonesia. Because we consider the Paratethyan taxon to be a subjective junior synonym of Oxymeris modesta , no replacement name is needed.

Terebra (Subula) plicaria perstriatula Sacco, 1891 ; holotype, GBA, SL: 67.5 mm, MD: 18.8 mm, Lăpugiu des Sus ( Romania). The specimen was stored in the Geosphere Austria (formerly Geological Survey of Austria) but is lost. This name was established by Sacco (1891a) for the specimen illustrated by Hoernes & Auinger (1880: pl. 12, fig. 19) as “ Terebra hochstetteri ??”. This fragmentary specimen represents an Oxymeris modesta with delicate spiral grooves on the last teleoconch whorls.

Discussion. The authorship of Oxymeris modesta is frequently given as Tristan in Defrance 1829, but as this was a manuscript name, as clearly stated by Defrance (1829: 288), the authorship must pass to Defrance (1829). Paratethyan material of this species was variously identified as Oxymeris plicaria (de Basterot, 1825) or O. modesta ( Defrance, 1829) in the literature. Oxymeris plicaria is based on material from the Burdigalian of Saucats ( France) ( Figs 12A–C View FIGURE 12 ) and O. modesta is based on specimens from the Langhian of the Touraine ( France) ( Fig. 13F View FIGURE 13 ). Bouchet (1981) assumed an anagenetic development from the Early Miocene Oxymeris plicaria (de Basterot, 1825) to the Middle and Late Miocene O. modesta ( Defrance, 1829) , which was said to be more obtuse, having a shallower suture and a less distinct subsutural groove ( Bouchet 1981: 365). We have studied Early Miocene specimens from Saucats ( France) (10 spec., NHMW 1851/0019/0005; 6 spec., NHMW 1850/0018/0105; 6 spec., NHMW 1856/0029/0028), Bordeaux ( France) (2 spec., NHMW 1836/0012/0185) Saint-Paul-lès-Dax ( France) (4 spec., 1850/0018/0024), Middle Miocene specimens from Salles ( France) (4 spec., NHMW1856/0035/0492) Sainte-Maure, Pontlevoy ( France) (12 spec., NHMW 1854/0018/0018) and Late Miocene specimens from Tortona ( Italy) (1 spec., NHMW 1862/0035/0215) and Cacela ( Portugal (3 spec., NHMW 1863/0057/0023). The comparison between these specimens showed that the shell of Oxymeris plicaria has more prominent axial ribs on early teleoconch whorls, persisting down to about 8–12 whorls ( Figs 12A–C View FIGURE 12 ). Oxymeris modesta differs in its larger maximum size (but small adult specimens occur as well), usually has weaker axial sculpture, which fades earlier around 8–9 th teleoconch whorls, has more convex whorls and is slightly more obtuse. The big confusion between both species was mainly caused by the fact that researchers such as Davoli (1977) identified Middle and Late Miocene specimens of O. modesta as O. plicaria . This confusion induced later authors, such as Bałuk (1997) and Landau et al. (2013) to treat both species as morphs of a single species (which was correct based on Davoli’s material). Herein, we follow Bouchet (1981) and consider the Early Miocene Oxymeris plicaria and the latest Early to Middle Miocene O. modesta as anagenetic succession of two (chrono)species. The Paratethyan specimens and those from the Serrvallian of the Karaman Basin ( Turkey) represent typical O. modesta .

Based on Pliocene material from Italy (e.g., Asti, 21 spec., NHMW 1851/0017/0052), Castell’Arquato, 5 spec., NHMW 1831/0017/0054), we agree with Landau et al. (2013) in separating Oxymeris fuscata ( Brocchi, 1814) from O. modesta due to its more elongate shell, with more flat sided whorls, weaker axial sculpture and narrower aperture. The Pliocene O. fuscata ( Brocchi 1814) was discussed by Bouchet (1981) to have entered the Mediterranean Sea from the Atlantic after the Messinian Salinity Crises.

Paleoenvironment. Ubiquitous in all paleoenvironments from coastal marine to outer neritic.

Distribution in Central Paratethys. Karpatian (Early Miocene): Korneuburg Basin: Karnabrunn, Niederkreuzstetten, Stetten ( Austria) ( Hoernes & Auinger 1880; Harzhauser 2002); North Alpine Foreland Basin: Neuruppersdorf ( Austria) (hop opus), Slup ( Czech Republic) (Tejkal et al. 1968). Badenian (Middle Miocene): North Alpine-Carpathian Foreland Basin: Grund ( Austria) ( Hörnes 1852), Hrušovany nad Jevišovkou ( Czech Republic) ( Hoernes & Auinger 1880); Korytnica Basin: Korytnica ( Poland) ( Bałuk 1997); Polish-Ukrainian Fore-Carpathian Basin: Zgłobice ( Poland), Żabiak, Żukowce (Zhukivtsi) Zaleśce (Salisze) ( Ukraine) ( Friedberg 1911, 1938); Voronyaky Hills: Hołubica (Holubytsia) ( Ukraine) ( Friedberg 1911); Vienna Basin: Bad Vöslau, Baden, Baden-Sooss, Enzesfeld, Gainfarn, Möllersdorf, Steinebrunn, Traiskirchen, Vienna /Pötzleinsdorf ( Austria) ( Hoernes & Auinger 1880), Mikulov, Borský Mikuláš ( Czech Republic) ( Hörnes 1852; Švagrovský 1982); Eisenstadt-Sopron Basin: Forchtenau ( Austria) (hoc opus); Oberpullendorf Basin: Ritzing ( Austria) ( Hörnes 1852); Styrian Basin: Pöls ( Austria) (hoc opus); Pannonian Basin: Hidas, Pécsszabolcs, Várpalota, Szob, Letkés ( Hungary) ( Csepreghy-Meznerics 1950; Bohn-Havas 1973); southern Pannonian Basin: Miljevići ( Bosnia and Herzegovina) ( Atanacković 1985). Bükk Mountains: Bóta ( Hungary) ( Csepreghy-Meznerics 1972); Cserhát Mountains: Mátraverebély, Sámsonháza ( Hungary) ( Csepreghy-Meznerics 1954); Făget Basin: Bujtur, CoŞteiu de Sus, Lăpugiu des Sus, NemeŞeŞti ( Romania) ( Boettger 1902; hoc opus); Caransebeş-Mehadia Basin: Valea Bela Reca, Valea Calvei ( Romania) ( Hinculov 1968); Şimleu Basin: Tusa ( Romania) ( Nicorici 1972); Dacian Basin: Târnene, Staropatica ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

North Sea. Late Burdigalian-Langhian (Early/Middle Miocene): North Sea Basin: Netherlands (A.W. Janssen 1984), Germany ( Kautsky 1925; Moths et al. 2010).

Northeastern Atlantic. Aquitanian and Burdigalian (Early Miocene): Aquitaine Basin ( France) ( Peyrot 1931; Lozouet et al. 2001); Langhian-Serravallian (Middle Miocene): Aquitaine Basin ( France) ( Peyrot 1931); Langhian: Loire Basin, France ( Glibert 1952a); Tortonian (Late Miocene): Algarve Basin: Cacela ( Portugal) ( Pereira da Costa 1866).

Proto-Mediterranean Sea. Burdigalian (Early Miocene): Colli Torinesi ( Italy) ( Sacco 1891a); Langhian (Middle Miocene): Kasaba Basin ( Turkey) ( İslamoğlu 2004); Serravallian (Middle Miocene): Karaman Basin, Turkey ( Erünal-Erentöz 1958; Landau et al. 2013); Tortonian and Messinian (Late Miocene): Po Basin ( Italy) ( Sacco 1891a; Davoli 1977).

NHMW

Naturhistorisches Museum, Wien

Kingdom

Animalia

Phylum

Mollusca

Class

Gastropoda

Order

Neogastropoda

Family

Terebridae

Genus

Oxymeris

Loc

Oxymeris modesta ( Defrance, 1829 )

Harzhauser, Mathias & Landau, Bernard M. 2023
2023
Loc

Terebra (Subula) fuscata var. modesta

Kojumdgieva, E. M. & Strachimirov, B. 1960: 218
1960
Loc

Terebra (Subula) fuscata (Brocchi)

Erunal-Erentoz, L. 1958: 125
1958
Loc

Terebra (Subula) plicaria Bast.

Csepreghy-Meznerics, I. 1954: 57
1954
Loc

Terebra (Subula) fuscata plicaria

Strausz, L. 1954: 37
1954
Loc

Terebra (Oxymeris) modesta

Glibert, M. 1952: 380
1952
Loc

Terebra (Subula) plicaria Bast.

Csepreghy-Meznerics, I. 1950: 62
1950
Loc

Terebra modesta

Friedberg, W. 1938: 158
1938
Loc

Terebra (Subula) modesta Tristan in Defrance

Peyrot, A. 1931: 121
1931
Loc

Terebra (Subula) fuscata Brocc.

Friedberg, W. 1911: 1
1911
Loc

Terebra (Acus) fuscata Brocc.

Boettger, O. 1902: 18
1902
Loc

Terebra (Subula) plicaria

Sacco, F. 1891: 14
1891
Loc

[Terebra] S [ubula]. plicaria var. laevisulcata

Sacco, F. 1891: 12
1891
Loc

[Terebra] Subula conicoplicaria

Sacco, F. 1891: 15
1891
Loc

[Terebra] Subula modesta ( Tristan

Sacco, F. 1891: 15
1891
Loc

Terebra] S [ubula]. modesta var. plano-clavata

Sacco, F. 1891: 16
1891
Loc

[Terebra] S [ubula]. modesta var. perinflata

Sacco, F. 1891: 16
1891
Loc

[Terebra] S [ubula]. modesta var. dertofusulata

Sacco, F. 1891: 16
1891
Loc

[Terebra] S [ubula]. modesta var. ovulata

Sacco, F. 1891: 16
1891
Loc

Terebra (Acus)

Hoernes, R. & Auinger, M. 1880: 107
1880
Loc

Terebra Hochstetteri

Hoernes, R. & Auinger, M. 1880: 107
1880
Loc

Terebra fuscata Brocc.

Pereira da Costa, F. A. 1866: 78
1866
Loc

Terebra Blainvillei

Eichwald, E. 1853: 161
1853
Loc

Terebra fuscata Brocc.

Hornes, M. 1852: 128
1852
Loc

Terebra Blainvillei

Eichwald, E. 1851: 89
1851
Loc

Terebra fuscata

Hornes, M. 1848: 17
1848
Loc

Terebra fuscata

Hauer, J. 1837: 417
1837
Loc

Subula

Eichwald, E. 1830: 223
1830
Loc

Terebra modesta

Defrance, M. 1829: 288
1829
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