Lasius tapinomoides , Salata, Sebastian & Borowiec, Lech, 2018

Salata, Sebastian & Borowiec, Lech, 2018, A new species of the ant genus Lasius Fabricius, 1804 from Crete (Hymenoptera, Formicidae), ZooKeys 789, pp. 139-159: 142-153

publication ID

http://dx.doi.org/10.3897/zookeys.789.27022

publication LSID

lsid:zoobank.org:pub:0E40808C-D9E6-43D6-AA8E-D17DD66B63A8

persistent identifier

http://treatment.plazi.org/id/412DD1F4-21DA-4A85-B4CB-5031FC049560

taxon LSID

lsid:zoobank.org:act:412DD1F4-21DA-4A85-B4CB-5031FC049560

treatment provided by

ZooKeys by Pensoft

scientific name

Lasius tapinomoides
status

sp. n.

Lasius tapinomoides  sp. n. Figs 1-2, 3-5, 6-7

Type material.

Holotype (w.): ‘‘ Lasius  / tapinomoides  sp. nov. / HOLOTYPE // Collection L. Borowiec / Formicidae  / LBC-GR00976 // GREECE, Crete, Rethymno Pr. / Antonios Spilia Gorge / 35° 15.245 N,24° 34.220 E / 11 V 2013, 342 m / L.Borowiec // CASENT0845075 ’’ ( DBET); paratypes (6w.,1q.): data same as holotype, CASENT0845076 to CASENT0845082 ( DBET, NHMC, NHMB); paratypes (15w.), CASENT0845460 to CASENT0845474: ‘‘ GREECE, Crete, Rethymno Pr. / Kato Malaki / 35.28333 N, 24.4 E / 15 V 2013, 235 m / L. Borowiec’’ ( DBET, NHMC, BMNH).

Non-type material.

2w. (pin): ‘‘ Collection L. Borowiec / Formicidae  / LBC-GR00467 // GREECE, W Crete, 339 m / Plemeniana n. Kandanos / 35.31666 N, 23.71666 E / 2 V 2011, L. Borowiec ( DBET); 1w. (pin): Collection L. Borowiec / Formicidae  / LBC-GR01420 // GREECE, Crete, Rethymno / Orthes Gorge, 318 m / 35.3336 N, 24.6848 E / 28 IV 2014, S. Salata’’ ( DBET); 2w. (pin): ‘‘ Collection L. Borowiec / Formicidae  / LBC-GR00993 // GREECE, Crete, Rethymno / n. Argioupolis / 35.28333 N, 24.33333 E / 13 V 2013, 197 m / L. Borowiec’’ ( DBET); 1w. (pin): ‘‘ GREECE, Crete, Rethymno Pr. / Preveli Beach / 35.16666 N, 24.46666 E / 7 V 2013, 10 m / L. Borowiec’’ ( DBET); 7w. (pin): ‘‘ GREECE, Crete, Rethymno Pr. / Road to Preveli Beach loc. 1 / 35.16666 N, 24.45 E / 7 V 2013, 58 m / L. Borowiec’’ ( DBET; 12w. (EtOH), Crete, Lasithi Prov., Gorge of Richtis, 35.16667N, 25.98333E, 245m, 08.iv.2014, leg. S. Salata ( DBET); 2w. (EtOH), Orthes Gorge, 35.3336N, 24.6848E, 318 m, 28.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Antonios Spilia Gorge, 35° 15.245 N,24°34.220, 342m, 11.v.2013, leg. S. Salata ( DBET); 5w. (EtOH), road to Preveli Beach loc. 1, 35.16666N, 24.45E, 58m, 7.v.2013, leg. S. Salata ( DBET).

Etymology.

The name refers to the similarity of this species to species of the Tapinoma  genus, caused by a very shallow metanotal groove.

Description.

Worker.

Measurements: see Table 1.

Head, mesosoma, petiole and gaster uniformly coloured, brown to dark brown. Antennae, tibiae and tarsi bright brown to orange (Figs 1-4).

Head oval, 1.2 times as wide as long, lateral surfaces above eyes convex, occipital margin of head slightly convex (Figs 3-4). Clypeus shiny and smooth, its anterior margin convex, lacking median anterior notch, covered with sparse, decumbent to erect pubescence, average distance between setae longer than three fourths of their length. Masticatory border of mandibles with 7-8 teeth. Eyes medium-sized, oval, 0.25 times as long as length of the head. Antennal scape long, straight or slightly curved on its anterior part, 0.9 times as long as length of the head, exceeding beyond occipital margin of head, in apex gradually widened. Pedicel more than 2.0 times longer than wide, average 2.5 times longer than second segment of funiculus. Other funicular segments from 1.5 to 2.0 times longer than wide (Figs 3-4). Surface of scape with very fine microreticulation, shiny. Its surface covered with thin, dense, adpressed setae, on its apical part several suberect setae also occur.

Genae with few adpressed to suberect setae (Figure 3). Underside of head with thin, dense, adpressed setae and a few long, suberect to erect setae (Figure 1). Whole frontal head surface covered with short, adpressed and dense pubescence and sparse, long, thick suberect to erect setae, the distance between setae at least as long as three fourths of their length (Figs 3-4).

Mesosoma short, 1.9 times as long as wide. In lateral view, promesonotum low and flattened, metanotal groove very shallow, propodeum very low, propodeal dorsum slightly convex, propodeal declivity convex, less than twice length of propodeal dorsum (Figure 1). Whole mesosoma surface shiny with sparse microreticulation, covered with short, adpressed and dense pubescence, and sparse, long, thick suberect to erect setae, the distance between setae at least as long as half of their length, metapleuron below the level of the propodeal spiracle with more than 5 setae (Figs 1-2).

Petiole scale low, in lateral view with slightly convex sides, its dorsal crest thick and arched (Figure 1). Gaster with very rare and fine microreticulation, shiny, bearing pilosity similar to this covering mesosoma. Legs long, shiny, with fine microreticulation. Surface of tibia and femora with thin, dense, adpressed to subdecumbent setae, extensor profile without erect setae (Figs 1-2).

Description.

Gyne.

Measurements and Indices (n=1): HL: 1.03; HW: 1.2; SL: 0.9; EL: 0.3; EW: 0.24; ML: 2.3; HTL: 1.3; PNW: 1.4; HI: 116.5; SI1: 87.4; SI2: 75; MI: 56.5; EI1: 80; EI2:23; TI: 92.

Head, mesosoma, petiole and gaster dark brown. Antennae, tibiae and tarsi bright brown to orange (Figs 5-7).

Head trapezoidal, 1.1 times as wide as long, lateral surfaces above eyes convex, sides of occipital margin of head slightly convex, its central part concave (Figure 5). Clypeus shiny and smooth, its anterior margin convex, lacking median anterior notch, covered with decumbent to erect setae, average distance between setae longer than one third of their length. Masticatory border of mandibles with seven teeth. Eyes medium-sized, oval, 0.3 times as long as length of the head. Antennal scape short, curved on its anterior part, 0.9 times as long as length of the head, slightly exceeding beyond occipital margin of head, in apex gradually widened. Pedicel more than 2.0 times longer than wide, 2.0 times longer than second segment of funiculus. Other funicular segments from 1.5 to 2.0 times longer than wide (Figure 5). Surface of scape with very fine microreticulation, shiny. Its surface covered with thin, dense, adpressed to decumbent pubescence, on its apical part a few short, suberect setae also occur. Genae with thin, dense adpressed pubescence and a few suberect setae (Figure 5). Underside of head with thin, dense, adpressed pubescence and a few long, suberect setae (Figure 6). Whole frontal head surface covered with short, adpressed and dense pubescence and sparse, long, thick suberect to erect setae, the distance between setae at least as long as three fourths of their length (Figs 5-6).

Mesosoma long, 1.6 times as long as wide. In lateral view moderately high, its dorsum slightly convex, propodeal dorsum slightly convex, propodeal declivity convex (Figure 6). Whole mesosoma surface shiny with very sparse microreticulation, covered with short, adpressed and dense pubescence, and sparse, long, thick suberect to erect setae, the distance between setae at least as long as half of their length (Figs 6-7).

Petiole scale low and wide, in lateral view with slightly convex sides, its dorsal crest wide and deeply concave in central part. Gaster with moderately thick and fine microreticulation, shiny, bearing pilosity denser than this covering mesosoma. Legs long, shiny, with fine microreticulation. Surface of tibia and femora with thin, dense, adpressed to subdecumbent setae, extensor profile of tibia with erect setae (Figs 6-7).

Differential diagnosis.

Worker. As a member of the L. alienus  group it is characterized by dorsal plane of scape, genae, and extensor profile of hind tibiae lacking or having very few erect or suberect setae and, in all species known from Crete, presence of >15 erect setae on the occipital edge of the head. Within the L. alienus  group it can be classified to the L. turcicus  complex. This complex can be characterized by small number of mandibular teeth (6-8), usually lack of suberect setae on hind tibia, very sparse clypeal pubescence, and more or less shallow metanotal groove. There are three known species of this complex: L. turcicus  , L. neglectus  and L. austriacus  . Lasius tapinomoides  sp. n. differs from all of them in presence of suberect to erect setae on antennal scape covering its apical part (ca. 1/3 upper part of the scape). Additionally from first two relatives it differs also in very shallow metanotal groove and from L. austriacus  it differs in more flattened promesonotum, antennal sockets set not very close to posterior clypeal margin and habitat preferences. Lasius austriacus  is related with xerothermous sites ( Schlick-Steiner et al. 2003, Steiner et al. 2004) while L. tapinomoides  inhabits moist, closed canopy forests.

There are two other species of the L. alienus  group known from Crete: L. bombycina  Seifert & Galkowski, 2016 and L. turcicus  Santschi, 1921. Lasius tapinomoides  differs from all of them in very small body size. Nevertheless, at the first glance it can be confused with small workers of L. turcicus  , from which it differs in the following measurements ( L. tapinomoides  sp. n. vs L. turcicus  ): HI: 81.8 ± 3.3 (73.4-85.0) vs 90.2 ± 1.8 (87.2-92.8), SI2: 115.7 ± 4.4 (112.2-127.6) vs 104.7 ± 3.6 (97.3-109.3), TI: 82.0 ± 2.6 (75.8-85.2) vs TI: 89.2 ± 2.2 (85.4-93.8). For more measurements data see Table 1.

Gyne. During our fieldwork we could observe several gynes of L. tapinomoides  and all detected nests were monogynous. Unfortunately, we were able to collect only a single specimen, therefore we provide very scarce data. As a coherent differential diagnosis is impeded, we decided to limit it to the most visible difference. Based on the morphology, L. tapinomoides  differs from L. turcicus  , L. austriacus  and L. neglectus  in presence of erect setae on tibiae.

General distribution.

Greece: Crete - endemic species.

Biology.

Species inhabiting moist, closed canopy forests, which are most often located in stream valleys. Nesting in wet soil, under shallow and small rocks. Nests, most often, located in the vicinity of water sources. Workers were found in the litter or on the rocks surrounding the nest entrance. Colonies monogynous.

The following ant species were recorded in the same areas as L. tapinomoides  :

Antonios Spilia Gorge: Aphaenogaster cecconii  Emery, A. rugosoferruginea  Forel, A. simonellii  Emery, Camponotus candiotes  Emery, C. lateralis  (Olivier), Crematogaster ionia  Forel, Messor wasmanni  Krausse, Pheidole cf. pallidula  , Stigmatomma denticulatum  Roger, Temnothorax ariadnae  Csősz, Heinze & Mikó, T. cf. graecus  , T. cf. exilis  , T. cf. luteus  , Tetramorium caespitum  (Linnaeus), T. diomedeum  Emery;

Plemeniana n. Kandanos: Aphaenogaster rugosoferruginea  , A. simonellii  , Camponotus candiotes  , C. gestroi  Emery, C. lateralis  , Colobopsis truncata  (Spinola), Crematogaster cf. ionia  , Lasius lasioides  (Emery), Messor ibericus  Santschi, Tetramorium caespitum  ;

Orthes Gorge: Aphaenogaster cecconii  , A. cf. subterranea  , A. simonellii  , Camponotus baldaccii  Emery, C. candiotes  , C. jaliensis  Dalla Torre, C. kiesenwetteri  (Roger), C. rebeccae  Forel, Crematogaster sordidula  (Nylander), Lasius lasioides  , Lepisiota nigra  Dalla Torre, Messor cf. muticus  , M. wasmanni  , Tetramorium cf. caespitum  , T. kephalosi  Salata & Borowiec

n. Argioupolis: Aphaenogaster cecconii  , A. rugosoferruginea  , A. simonellii  , Camponotus candiotes  , C. lateralis  , Crematogaster cf. ionia  , Lepisiota nigra  , Messor cf. muticus  , M. wasmanni  , Pheidole cf. pallidula  , Plagiolepis pallescens  sensu Radchenko, Temnothorax cf. luteus  , T. ariadnae  , T. cf. exilis  , T. cf. graecus  , Tetramorium cf. caespitum  ;

Preveli Beach: Aphaenogaster cecconii  , Crematogaster ionia  , Lasius psammophilus  Seifert, Lepisiota melas  (Emery), Pheidole cf. pallidula  , Tapinoma festae  Emery, Tetramorium kephalosi  ;

road to Preveli Beach loc. 1: Aphaenogaster rugosoferruginea  , A. simonellii  , Camponotus baldaccii  , C. candiotes  , C. gestroi  , C. kiesenwetteri  , C. lateralis, Cardiocondyla mauritanica  Forel, Crematogaster cf. ionia  , Cryptopone ochracea  (Mayr), Lasius lasioides  , L. bombycina  Seifert & Galkowski, L. psammophilus  , Lepisiota nigra  , Messor ibericus  , M. wasmanni  , Pheidole cf. pallidula  , Plagiolepis pallescens  sensu Radchenko, Temnothorax cf. luteus  , T. cf. graecus  , T. recedens  (Nylander), Tetramorium cf. caespitum  ;

Kato Malaki: Aphaenogaster simonellii  , Camponotus baldaccii  , C. gestroi  , C. jaliensis  , Crematogaster cf. ionia  , Crematogaster sordidula  , Messor ibericus  , M. wasmanni  , Pheidole cf. pallidula  , Plagiolepis pallescens  sensu Radchenko, Temnothorax ariadnae  , T. recedens  , Tetramorium diomedeum  , Tetramorium punctatum  Santschi;

Gorge of Richtis: Aphaenogaster rugosoferruginea  , Camponotus candiotes  , C. gestroi  , C. lateralis  , Crematogaster sordidula  , Lasius lasioides  , Messor wasmanni  , Temnothorax cf. exilis  , T. cf. graecus  , Tetramorium cf. caespitum  .

Comments.

We examined several hundred specimens of Lasius turcicus  from 76 samples across Greece and western Turkey (including initial nest samples), also numerous samples from Crete (see new material listed below). In none of these samples did we find specimens with such a shallow metanotal groove like in L. tapinomoides  . This character is constant in all examined specimens and is always correlated with very small body size of workers and preference to humid habitats. Across all the sampled area within Greece and Turkey, where we collected species of the L. turcicus  complex, we have not found workers or nests with workers similar to L. tapinomoides  . Moreover, L. tapinomoides  is the only known member of the L. turcicus  complex that have suberect to erect setae on the apical 1/3 part of the antennal scape. This has prompted us to hypothesize that Cretan samples represent a new species.

New records of Cretan members of the genus Lasius 

Lasius bombycina  Seifert & Galkowski, 2016

GREECE, Crete: 2w (EtOH), Omalos, 35.31667N, 23.9E, 1122 m, 03.v.2014, leg. S. Salata ( DBET); 4w. (pin), 03.v.2011, Omalos Plateau, 35.33333N, 23.88333E, 1034 m, leg. L. Borowiec ( DBET); 1w. (EtOH), Potamida  n. Mythimna, 35.46666N, 23.68333E, 37 m, 02.v.2011, leg. L. Borowiec ( DBET); 1w. (pin), W of Georgioupoli, 35.36666N, 24.25E, 17 m, 02.v.2007, leg. L. Borowiec ( DBET); 8w. (EtOH), Zaros Lake, 35.13333N, 24.9E, 409 m, 26.iv.2014, leg. S. Salata (DBET); 3w. (EtOH), Kato Symi loc. 3, 35.05N, 25.48333E, 818 m, 16.iv.2014, leg. S. Salata ( DBET); 4w. (EtOH), Lasithi Platou - Plati, 35.16667N, 25.43333E, 831 m, 01.v.2014, leg. S. Salata ( DBET); 2w. (EtOH), Orino, 35.06667N, 25.9E, 523 m, 12.iv.2014, leg. S. Salata ( DBET); 2w. (pin), Ag. Joannis Forest loc. 1, 35.23333N, 24.4E, 448 m, 06.v.2013, leg. L. Borowiec ( DBET); 1w. (pin), Ampelakiou, 35.26666N, 24.46666E, 464 m, 10.v.2013, leg. L. Borowiec ( DBET); 1w. (EtOH), Fourfouras, 35.21666N, 24.71666E, 578 m, 14.v.2013, leg. S. Salata ( DBET); 5w. (EtOH), n. Velonado, 35.25N, 24.36667E, 373 m, 13.v.2013, leg. S. Salata ( DBET); 2w. (EtOH), n. Vilandredo, 35.25N, 24.31667E, 354 m, 13.v.2013, leg. S. Salata ( DBET); 2w. (EtOH), road to Preveli Beach loc. 1, 35.16666N, 24.45E, 58 m, 7.v.2013, leg. S. Salata ( DBET); 3w. (EtOH), Selli-Oros rd., 35.28333N, 24.5E, 473 m, 11.v.2013, leg. S. Salata, ( DBET); 1w. (pin), Spili-Gerakari rd. loc. 2, 35.21987N, 24.57144E, 804 m, 09.v.2013, leg. L. Borowiec ( DBET); 1w. (EtOH), Asi Gonia, 35.25N, 24.26667E, 716 m, 29.v.2001, leg. S. Salata ( DBET); 1w. (EtOH), Niato plateau, 35.28333N, 24.13333E, 1200 m, 19.vii.2013, leg. S. Salata ( DBET); 1w. (EtOH), Therisso to Kaloros Mt., 35.35N, 23.95E, 1130 m., 31.xii.2013, leg. Simaiakis ( NHMC); 1w. (EtOH), Therisso to Kaloros Mt., 35.3667N, 23.9833E, 1134 m., 19.Vii.2013, leg. Simaiakis ( NHMC); 1w. (EtOH), Diplori, 35.1667N, 24.9333E, 1350 m., 19.x.1999, leg. Nikolakakis ( NHMC); 1w. (EtOH), Dikti Mt., 35.11667N, 25.4667E, 1450 m., 9.i.2001, leg. Simaiakis ( NHMC); 1w. (EtOH), Tigania, 35.28333N, 24.7333E, 1100 m., 7.iv.2000, leg. Nikolakakis ( NHMC).

Lasius myops  Forel, 1894

GREECE, Crete: 3w. (pin), 3w. (EtOH), Anopoli, 35.26667N, 24.06667E, 1780 m, 22.vii.2006, leg. Chatzaki M. ( NHMC, DBET).

Lasius illyricus  Zimmermann, 1935

GREECE, Crete: 6w. (pin), Limnakarou platou, 35.13333N, 25.46667E, 1130 m, 26.iv.2014, leg. S. Salata ( DBET); 6w. (EtOH), Limnakarou platou, 35.13333N, 25.46667E, 1130 m, 10.iv.2014, leg. S. Salata ( DBET); 2w. (EtOH), Dikti mt., 35.1N, 25.46667E, 1750 m, 05.viii.2000, leg. M. Chatzaki ( NHMC).

Lasius lasioides  (Emery, 1869)

GREECE, Crete: 1w. (pin), Agia, 6 km SW Chania, 35.46666N, 23.91666E, 22 m, 03.v.2011, leg. L. Borowiec ( DBET); 5w. (pin), Askifou, 35.26666N, 24.16666E, 730 m, 01.v.2007, leg. L. Borowiec & M.L. Borowiec ( DBET); 7w. (EtOH), Diktamos Gorge n. Stilos, 35.43333N, 24.1E, 160 m, 04.v.2011, leg. L. Borowiec ( DBET); 3w. (EtOH), Kalives river, 35.45N, 24.13333E, 26 m, 01.v.2014, leg. S. Salata ( DBET); 1w. (pin), Kourna Lake, 35.31666N, 24.28333E, 95 m, 03.v.2007, leg. L. Borowiec & M.L. Borowiec ( DBET); 1w. (pin), Koutsomatados-Mili rd., 35.38333N, 23.66666E, 308 m, 02.v.2011, leg. L. Borowiec ( DBET); 2w. (EtOH), Omalos, 35.31667N, 23.9E, 1122 m, 03.v.2014, leg. S. Salata ( DBET); 1w. (pin), Plemeniana n. Kandanos, 35.31666N, 23.71666E, 339 m, 02.v.2011, leg. L. Borowiec ( DBET); 4w. (EtOH), Potamida  n. Mythimna, 35.46666N, 23.68333E, 37 m, 02.v.2011, leg. L. Borowiec ( DBET); 17w. (EtOH), Therisso Gorge, 35.43333N, 23.98333E, 320 m, 01.v.2011, leg. L. Borowiec ( DBET); 3w. (EtOH), Ganies–Kalamaki road, 35.28333N, 24.93333E, 439 m, 16.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Iraklion city-walls, 35.31667N, 25.11667N, 46 m, 03.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Kastelli 1 km E, 35.2N,25,33333E, 363 m, 15.iv.2014, leg. S. Salata ( DBET); 6w. (EtOH), Katofigi, 35.08333N, 25.4E, 560 m, 12.iv.2014, leg. S. Salata ( DBET); 2w. (EtOH), Miamou, 34.96667N, 24.93333E, 494 m, 24.iv.2014, leg. S. Salata ( DBET); 2w. (EtOH), Rouvas Forest loc. 1, 35.15N,24.93333, 1316 m, 10.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Rouvas Gorge, 35.13333N, 24.9E, 455 m, 26.iv.2014, leg. S. Salata ( DBET); 5w. (EtOH), Xanias–Miliarades road, 35.08333N, 25.38333E, 504 m, 13.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Dead’s Gorge, 35.08333N, 26.25E, 15 m, 09.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Gorge of Richtis, 35.16667N, 25.98333E, 245 m, 08.iv.2014, leg. S. Salata ( DBET); 4w. (EtOH), Hristos–Mathokotsana road, 35.08333N, 25.56667E, 703 m, 11.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Kalami–Psari Forada road, 35.016667N,25.48333, 419 m, 12.iv.2014, leg. S. Salata ( DBET); 5w. (EtOH), Kato Symi loc. 1, 35.05N, 25.48333E, 1206 m, 12.iv.2014, leg. S. Salata ( DBET); 5w. (EtOH), Lasithi Platou - Pinakiano, 35.18333N, 25.45E, 806 m, 23.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Lastros, 35.13333N, 25.88333E, 336 m, 06.iv.2014, leg. S. Salata ( DBET); 10w. (EtOH), Limnakarou plato, 35.13333N, 25.46667E, 1130 m, 26.iv.2014, leg. S. Salata ( DBET); 1w. (EtOH), Merisini  , 35.16667N, 25.9333E, 309 m, 06.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Mesa Lasithi, 35.16667N, 25.5E, 838 m, 28.iv.2014, leg. S. Salata ( DBET); 2w. (EtOH), Moni Kapsa, 35.01667N, 26.05E, 1 m, 10.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Neapoli–Vrisses road, 35.2333N, 25.6E, 443 m, 05.iv.2014, leg. S. Salata ( DBET); 2w. (EtOH), Perma–Koutounari road, 35.01667N, 25.83333E, 0 m, 11.iv.2014, S. Salata ( DBET); 2w. (EtOH), Schinokapsala–Agios Ioannis road, 35.05N,25.85, 452 m, 10.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Tourloti– Mirsini  road, 35.15N, 25.93333E, 266 m, 06.iv.2014, leg. S. Salata ( DBET); 2w. (EtOH), Voila, 35.08333N, 26.1E, 578 m, 10.iv.2014, leg. S. Salata ( DBET); 12w. (EtOH), Ag. Joannis Forest loc. 2, 35.23333N, 24.4E, 480 m, 06.v.2013, leg. S. Salata ( DBET); 2w. (EtOH), Chromonastiri, 35.326944N, 24.510278E, 262 m, 10.v.2013, leg. S. Salata ( DBET); 1w. (EtOH), Katsifou Gorge, 35.2N, 24.38333E, 57 m, 05.v.2013, leg. S. Salata ( DBET); 5w. (EtOH), Kissos, 35.18333N, 24.56667E, 623 m, 09.v.2013, leg. L. Borowiec ( DBET); 1w. (pin), Klisidi, 35.26666N, 24.63333E, 642 m, 06.v.2013, leg. L. Borowiec ( DBET); 4w. (EtOH), n. Velonado, 35.25N, 24.36667E, 373 m, 13.v.2013, leg. S. Salata ( DBET); 1w. (EtOH), n. Vilandredo, 35.25N, 24.31667E, 354 m, 13.v.2013, leg. S. Salata ( DBET); 3w. (EtOH), Orthes Gorge, 35.33333N,24.68333, 318 m, 28.iv.2014, leg. S. Salata ( DBET); 2w. (EtOH), Plakias, 35.191389N, 24.395E, 4 m, 05.v.2013, leg. S. Salata ( DBET); 41w. (EtOH), Plakias; Akrotiri Kakomouri, 35.16667N, 24.398055E, 28 m, 05.v.2013, leg. S. Salata ( DBET); 1w. (pin), road to Preveli Beach loc. 1, 35.16666N, 24.45E, 58 m, 07.v.2013, leg. L. Borowiec ( DBET); 1w. (pin), Spili, 35.21666N, 24.53333E, 537 m, 09.v.2013, leg. L. Borowiec ( DBET); 1w. (pin), Vistagi, 35.23333N, 24.68333E, 563 m, 06.v.2013, leg. L. Borowiec ( DBET); 1w. (pin), Xirokambos, 35.110556N, 24.558889E, 24 m, 12.v.2013, leg. L. Borowiec ( DBET); 3w. (EtOH), Kournas lake, 35.31667N, 24.26667E, 30 m, 10.vii.1997, leg. P. Lymberakis ( NHMC); 1w. (EtOH), Kardaki, 650 m, 35.2N, 24.61667E, 21.vii.1999, leg. E. Nikolakakis ( NHMC).

Lasius psammophilus  Seifert, 1992

GREECE, Crete: 4w. (EtOH), Agia Triada n. Kalamaki, 35.0508N, 25.7542E, 1 m, 28.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Moni Kapsa, 35.01667N, 26.05E, 1 m, 24.iv.2014, leg. S. Salata ( DBET); 2w. (EtOH), Episkopi beach, 35.35N, 24.35E, 0 m, 09.iv.2014, leg. S. Salata ( DBET); 1w. (pin), Gerakari, 35.21666N, 24.58333E, 751 m, 09.v.2013, leg. L. Borowiec ( DBET); 1w. (pin), Plakias, 35.191389N, 24.395E, 4 m, 05.v.2013, leg. L. Borowiec ( DBET); 4w. (pin), Preveli Beach, 35.154444N, 24.472778E, 10 m, 07.v.2013, leg. L. Borowiec ( DBET); 1w. (EtOH), Irakleio, 35.33333N, 25.13333E, 10 m, 18.ix.2010, leg. E. Panagiotou ( NHMC); 2w. (EtOH), Chamaitoulo, 35.03333N, 26.2E, 06.v.2001, 180 m, leg. E. Nikolakakis ( NHMC).

Lasius turcicus  Santschi, 1921

GREECE, Crete: 3w. (EtOH), Kalives river, 35.45N, 24.13333E, 26 m, 03.v.2014, leg. S. Salata ( DBET); 13w. (EtOH), Kato Daratso n. Chania, 35.5N, 23.983333E, 35-40 m, 07.v.2011, leg. L. Borowiec ( DBET); 4w. (pin), Koutsomatados–Mili rd., 35.38333N, 23.66666E, 308 m, 02.v.2011, leg. L. Borowiec ( DBET); 1w. (pin), Potamida  n. Mythimna, 35.46666N, 23.68333E, 37 m, 02.v.2011, leg. L. Borowiec ( DBET); 1w. (pin), Agia Triada n. Kalamaki, 35.0508N, 25.7542E, 1 m, 24.iv.2014, leg. S. Salata ( DBET); 2w. (EtOH), Agios Eirini, 35.26667N, 25.15E, 130 m, 02.iv.2014, leg. S. Salata ( DBET); 4w. (EtOH), Avgeniki, 35.18333N, 25.01667E, 227 m, 03.v.2014, leg. S. Salata ( DBET); 3w. (EtOH), Kastelli 1 km E, 35.2N, 25.33333E, 363 m, 16.iv.2014, leg. S. Salata ( DBET); 5w. (EtOH), Katofigi, 35.08333N, 25.4E, 560 m, 13.iv.2014, leg. S. Salata ( DBET); 2w. (EtOH), Panastros, 35.11667N, 24.98333E, 545 m, 03.v.2014, leg. S. Salata ( DBET); 5w. (EtOH), Sfendili, 35.25N, 25.38333E, 151 m, 06.iv.2014, leg. S. Salata ( DBET); 2w. (EtOH), Stoli–Louves road, 35.03333N, 25.01667E, 197 m, 30.iii.2014, leg. S. Salata ( DBET); 2w. (EtOH), Xanias–Miliarades road, 35.08333N, 25.38333E, 504 m, 25.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Dead’s Gorge, 35.08333N, 26.25E, 15 m, 09.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Hametoulo, 35.05N, 26.18333E, 520 m, 12.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Kalamafka, 35.06667N, 25.65E, 472 m, 12.iv.2014, leg. S. Salata ( DBET); 6w. (EtOH), Lasithi Platou - Pinakiano, 35.18333 N, 25.45E, 806 m, 28.iv.2014, leg. S. Salata ( DBET); 2w. (EtOH), Praisos, 35.11667N, 26.06667E, 193 m, 11.iv.2014, leg. S. Salata ( DBET); 6w. (EtOH), Ag. Joannis forest loc. 1, 35.23333N, 24.4E, 448 m, 06.v.2013, leg. S. Salata ( DBET); 2w. (EtOH), Chromonastiri, 35.326944N, 24.510278E, 262 m, 10.v.2013, leg. S. Salata ( DBET); 3w. (EtOH), Frati, 35.2N, 24.46666E, 297 m, 7.v.2013, leg. S. Salata ( DBET); 2w. (EtOH), Kissos, 35.18333N, 24.56667E, 623 m, 09.v.2013, leg. S. Salata ( DBET); 1w. (EtOH), Kissou Kambros, 35.16666N, 24.55E, 514 m, 14.v.2013, leg. S. Salata ( DBET); 3w. (EtOH), Palelimnos, 35.3N, 24.41666E, 262 m, 15.v.2013, leg. S. Salata ( DBET); 4w. (EtOH), Plakias, 35.191389N, 24.395E, 4 m, 05.v.2013, leg. S. Salata ( DBET); 5w. (EtOH), road to Nida platou, 35.25N, 24.88333E, 1166 m, 01.iv.2014, leg. S. Salata ( DBET); 3w. (EtOH), Setoures, 35.26667N, 24.38333E, 305 m, 15.v.2013, leg. S. Salata ( DBET); 2w. (EtOH), Vistagi, 35.23333N, 24.68333E, 563 m, 16.v.2013, leg. S. Salata ( DBET); 1w. (EtOH), Agia lake, 35.46667N, 23.93333E, 40 m, 08.vii.1997, leg. P. Lymberakis ( NHMC); 2w. (EtOH), Almyros river, 35.33469N, 25.05441E, 297 m, 03.vi.2012, leg. E. Aspradaki ( NHMC); 1w. (EtOH), Aposelemis, 35.33333N, 25.33333E, 7 m, 02.viii.2000, leg. A. Trichas ( NHMC); 1w. (EtOH), Dikti mt., 35.11667N,25.46667, 1450 m, 05.viii.2000, leg. S. Simaiakis ( NHMC); 2w. (EtOH), Dikti mt., 35.1N, 25.46667E, 1750 m, 10.v.2001, leg. M. Chatzaki ( NHMC); 2w. (EtOH), Before Amygdali after Neapoli, 35.2N, 25.58333E, 561 m, 06.viii.1997, leg. I. Stathi ( NHMC); 1w. (EtOH), Agios Titos, 35.18333N, 24.75E, 1000 m, 14.iv.2000, leg. M. Papadimitrakis ( NHMC); 1w. (EtOH), Ano Meros, 35.16667N, 24.65E, 750 m, 15.iv.2000, leg. I. Stathi ( NHMC); 3w. (EtOH), Garazo, 35.33333N, 24.78333E, 100 m, 07.iv.2000, leg. E. Nikolakakis ( NHMC); 1w. (EtOH), Tigania, 35.28333N,24.73333, 1100 m, 07.iv.2000, leg. E. Nikolakakis ( NHMC).

Key to Cretan Lasius  species (based on the worker caste)

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hymenoptera

Family

Formicidae

Genus

Lasius