Trichadenotecnum maculatum, Yoshizawa & Aldrete & Mockford, 2008

Yoshizawa, Kazunori, Aldrete, Alfonso Neri García & Mockford, Edward L., 2008, Systematics and biogeography of the New World species of Trichadenotecnum Enderlein (Insecta: Psocodea: ‘ Psocoptera’: Psocidae), Zoological Journal of the Linnean Society 153 (4), pp. 651-723 : 687-689

TRICHADENOTECNUM MACULATUM SP. NOV.

Material examined: Holotype. Male. Mexico: Nuevo León, 5 km W of Ciénega, hacia Laguna de Sánchez , 1200 m, 25.v.1977, ANGA ( UNAM) . Paratypes. 1 female, same data as holotype ; 1 male, 1 female, Nuevo León, 10 mi E Iturbide, Hwy. 60, 13.vi.1962, J. Campbell ( ISU) .

Etymology: The specific epithet is taken from Latin macula (= spotted), indicating the heavily spotted forewing in this species.

Description: Male. Head. White in ground colour; vertical and orbital markings pale brown; coronal suture black; frontal suture very narrowly bordered with pale brown band dorsally; frons with two pairs of pale brown spots ventrally; eye black, IO/D = 1.0; ocelli white, ocellar field black; antennal socket bordered with dark brown band; postclypeus with c. eight rows of pale brown spots at middle, fused with each other ventrally forming a large blackish brown marking, ventro-lateral corner white, lateral margin of postclypeus brown, dorsal margin without marking; anteclypeus brown. Antenna pale brown, pedicel and scape darker. Mouth parts dark brown; third segment of maxillary palpus pale brown.

Thorax. Prothorax brown. Mesonotum white; anterior surface of scutum brown; anterior margin of lateral lobe of scutum black; posterior region of scutum with pair of brown markings at middle; scutellum white, median part and anterior margin brown; postnotum brown. Metanotum brown, with pair of whitish oval spots on posterior regions of lobes of scutum, lateral corner white. Meso- and metapleurites brown with longitudinal white band medially.

Legs. Coxae dark brown; femora brown except middle part of fore and middle femora white; tibiae pale brown, with darker portion near distal ends; tarsi brown.

Forewing ( Fig. 17F ). Densely covered with small spots. Spots in cell a 1 distinct, basal one smaller. Opposing spots in cell r apparent for anterior one but posterior one very small and less distinct. Proximal band broad, strongly constricted around fork of M + Cu, with elongate paler pigmented area in cell CuA. Distal band somewhat indistinct in cell r 5, band in cell r 3 distinct. Spot on roof of cell m 3 large. Submarginal spots diffuse; spot in cell r 5 larger than others and distinct; spots in cell m 1, m 2 and m 3 small, sometimes indistinct. Marginal clouds clear, darker along veins. Hindwing. Hyaline, extensively covered with pale brown spots, some spots fused with each other to form a large marking in cell CuP; veins brown.

Abdomen. White, each segment with blackish brown band. Terminalia. Clunial arm ( Fig. 25A ) long and narrow, slightly arched, forming pointed free process apically, antero-ventrally with fleshy lobe covered with fine setae. Anterior margin of 8th sternum ( Fig. 25C ) concave; fusion with hypandrium very narrow. Epiproct lobe ( Fig. 25B ) expanded dorsally, wider than long, dorsal margin slightly arched, with rounded projection at middle, posterior surface almost flattened. Paraproct ( Fig. 25A ): ventro-lateral band short, triangular; distal process broad. Hypandrium ( Fig. 25C ) with weakly rugose region laterally; left process sinuate, directed postero-externally at apex; right process of almost equal width from base to apex, projected outward, then strongly bent posterointernally near base, with few denticles at apex; median tongue narrowing medially, then broadened apically, apical margin almost straight, dorsal keel triangular. Phallosome ( Fig. 25D ): pseudoparameres widely separated, longer than wide, directed posteriorly, apical margin rounded or somewhat truncated. Phallobase widest at anterior two-fifths, then narrowing to pointed anterior end.

Length. B 2.6–2.9; FW 3.0–3.7; HW 2.2–2.8.

Female. Colour and general morphology almost as in male; eye IO/D = 1.9. Genitalia: Egg guide of subgenital plate ( Fig. 26A ) widened near distal end, postero-lateral corners rounded, apical margin only slightly arched; body of subgenital plate with pair of broad extensions of sclerite antero-medially. Dorsal valve of gonapophyses ( Fig. 26B ) with long distal process; posterior lobe of external valve strongly projected posteriorly. Internal plate as in Figure 26C .

Length. B 2.8–3.0; FW 3.2–3.8; HW 2.1–2.9.

Distribution: Mexico.

Figures

Figure 17. Forewings of Trichadenotecnum erwini (A), T. chiapense (B), T. quaesitum (C), T. quaesitellum (D), T. cerrosillae (E), T. maculatum (F), T. neoleonense (G), T. ericium (H), T. brevicornum (I), T. concinnum (J), T. acutum (K) and T. nicaraguense (L.).

Figure 25. Male terminalia structures of Trichadenotecnum maculatum, showing terminalia in lateral view (A), epiproct in posterior view (B), hypandrium in ventral view (C) and phallosome in ventral view (D).

Figure 26. Female genitalia of Trichadenotecnum maculatum, showing subgenital plate (A), gonapophyses (B) and internal plate (C) in ventral view.