Trichadenotecnum chiapense, Yoshizawa & Aldrete & Mockford, 2008

Yoshizawa, Kazunori, Aldrete, Alfonso Neri García & Mockford, Edward L., 2008, Systematics and biogeography of the New World species of Trichadenotecnum Enderlein (Insecta: Psocodea: ‘ Psocoptera’: Psocidae), Zoological Journal of the Linnean Society 153 (4), pp. 651-723 : 680-682

TRICHADENOTECNUM CHIAPENSE SP. NOV.

Material examined: Holotype. Male. Mexico: Chiapas , Apicpac, orilla de Presa Malpaso, 53 km N Ocozocoautla, 1.vii.1981, ANGA & L. B. Menchaca López ( UNAM) . Paratype. Mexico: 1 male (no. 96); 2 males, Veracruz, Los Tuxtlas, c. 25 mi NW of UNAM Field Station , 350 m, 22.xii.1984, E. L. Mockford et al. ( UNAM) . BELIZE: 1 male, Cayo District, Chiquibul Forest Reserve, Las Cuevas , 9–12.xi.1994, T. King & A. Howe ( UNAM) ; 1 male, same locality and collectors, 16–17.vi.1995 ( UNAM) ; 1 male, Cayo District, Chiquibul Forest Reserve, San Pastor , 10–13.xi.1994, T. King & A. Howe ; 1 male, 1 female, same locality and collectors, 23–26.iii.1995 ( UNAM) ; 1 male, 1 female, Cayo District, Chiquibul Forest Reserve , 23.iii.1995, T. King & A. Howe ( UNAM) ; 3 males, MT 11, 10–13.iv.1996, A. Howe & T. King ( UNAM) ; 2 males, 1 female, MT 14, 10–13.vi.1996, T. King & A. Howe ( UNAM) .

Etymology: The specific epithet is the patronymic of ‘Chiapas’, the state in southern Mexico where the holotype was collected.

Description: Male. Head. White in ground colour; vertical and orbital markings blackish brown; coronal suture black; frontal suture bordered with blackish brown broad band dorsally; frons with two pairs of blackish brown bands, median pair narrow and lateral pair broad; gena almost blackish brown except narrow white crescent dorsal region; eye black, IO/D = 1.1; ocelli white, ocellar field black; antennal socket bordered with broad blackish brown band; postclypeus with c. four rows of blackish brown spots dorsally, fused with each other ventrally forming a large, T-shaped blackish marking, ventro-lateral region white; anteclypeus brown. Antenna pale brown, pedicel and scape darker. Mouth parts brown; 4th segment of maxillary palpus darker.

Thorax. Prothorax blackish brown. Meso- and metathorax mostly blackish brown except membranous regions white; mesoscutum pale, anterior surface of anterior lobe blackish brown, mesopleuron with broad longitudinal white band medially; anterior margins of meso- and metascutum and around notal wing processes of meso- and metanotum blackish brown.

Legs. Pale brown; each tibia with darker portion near distal end; ventral surface of fore and middle femur with pair of blackish brown markings.

Forewing ( Fig. 17B ). Spots in cell a 1 subequal in size, basal one paler. Opposing spots in cell r large, triangular, narrowly separated from each other. Proximal band broad in cell CuP and posterior half of cell CuA, with large paler spot in cells CuA and CuP, strongly constricted just below fork of M + Cu, broadly interrupted just beyond fork of M + Cu. Median spots sparse, partly fused with each other. Distal band narrow. Spot on roof of cell m 3 broad. Submarginal spots apparent; spot in cell r 5 larger than others; spot in cell m 3 small and rather faint. Marginal clouds apparent. Hindwing. Hyaline; veins brown.

Abdomen. White, with irregular blackish brown markings, 6th to 8th venters blackish brown. Terminalia. Clunial arm ( Fig. 19A ) long and narrow, sicklelike in shape apically, its apical tip pointed forming free process. Eighth venter ( Fig. 19C ) with transverse sclerite, anterior margin almost straight, free from hypandrium posteriorly. Epiproct ( Fig. 19A, B ) chairshaped; epiproct lobe long and broad, broader than wide, slightly expanded dorsally, dorsal margin strongly sinuated, with small process at middle, posterior surface slightly swollen. Hypandrium ( Fig. 19C ): left process with flattened lateral surfaces, directed posteriorly, slightly sinuated, of almost equal width in lateral view from base to apical two-fifths then strongly narrowing to sharply pointed apex; right process much shorter than left process, lamellate, usually serrated apically; median tongue broadened apically, apical margin with or without shallow and broad notch, dorsal surface with semicircular keel covered by long spines. Phallosome ( Fig. 19D ): pseudoparameres widely separated, narrow, rounded apically, postero-internal part with group of denticles. Phallobase widest near middle, then narrowing to pointed anterior end.

Length. B 2.0–2.4; FW 2.6–2.9; HW 1.8–2.0.

Female. Colour and general morphology almost as in male; eye IO/D = 2.0. Genitalia: Egg guide of subgenital plate ( Fig. 20A ) relatively narrow, gradually narrowing apically, apical margin rounded; body of subgenital plate with pair of tongue-shaped extensions of sclerite antero-medially. Dorsal valve of gonapophyses ( Fig. 20B ) with long distal process; posterior lobe of external valve strongly projected posteriorly. Internal plate as in Figure 20C .

Length. B 2.5–2.8; FW 3.0–3.3; HW 2.1–2.4.

Distribution: Mexico, Belize.

Figures

Figure 17. Forewings of Trichadenotecnum erwini (A), T. chiapense (B), T. quaesitum (C), T. quaesitellum (D), T. cerrosillae (E), T. maculatum (F), T. neoleonense (G), T. ericium (H), T. brevicornum (I), T. concinnum (J), T. acutum (K) and T. nicaraguense (L.).

Figure 19. Male terminalia structures of Trichadenotecnum chiapense, showing terminalia in lateral view (A), epiproct in posterior view (B), hypandrium in ventral view (C) and phallosome in ventral view (D).

Figure 20. Female genitalia of Trichadenotecnum chiapense, showing subgenital plate (A), gonapophyses (B) and internal plate (C) in ventral view.