Pholcus khaolek Huber

Berhard A. Huber, Booppa Petchard, Charles Leh Moi Ung, Joseph K. H. Koh & Amir R. M. Ghazali, 2016, The Southeast Asian Pholcus halabala species group (Araneae, Pholcidae): new data from field observations and ultrastructure, European Journal of Taxonomy 190, pp. 1-55: 21-24

publication ID

10.5852/ejt.2016.190

publication LSID

lsid:zoobank.org:pub:urn:lsid:zoobank.org:pub:BE92596B-62D9-46CD-8486-CF6B36C640B11

persistent identifier

http://treatment.plazi.org/id/2A1CAF47-9EB4-480C-8C83-DFB0162AF617

taxon LSID

lsid:zoobank.org:act:2A1CAF47-9EB4-480C-8C83-DFB0162AF617

treatment provided by

Jeremy

scientific name

Pholcus khaolek Huber
status

sp. nov.

Pholcus khaolek Huber  , sp. nov.

urn:lsid:zoobank.org:act:2A1CAF47-9EB4-480C-8C83-DFB0162AF617

Figs 52–53View Figs 49 – 56, 63–78View Figs 63 – 67View Figs 68 – 78, 99–101View Figs 90 – 101

Diagnosis

Easily distinguished from most similar known relative ( Ph. kuhapimuk  sp. nov.) by morphology of male palps ( Figs 63–64View Figs 63 – 67; shapes of male palpal tarsus and procursus tip, larger uncus, complex appendix with retrolateral process) and by female internal genitalia ( Figs 66–67View Figs 63 – 67; distinctive median triangular sclerite). From other close relatives also by combination of pale coloration ( Figs 52–53View Figs 49 – 56), shape of male palpal trochanter apophysis (curved, proximally wide, distally pointed; Fig. 64View Figs 63 – 67), and shape and position of pore plates ( Fig. 67View Figs 63 – 67).

Etymology

The species name is derived from the type locality; noun in apposition.

Type material

THAILAND: holotype, ♂, ZFMK ( Ar 15028), Nakhon Si Thammarat, Khao Nan National Park, Tham Khao Lek (8°46.09' N, 98°43.68' E), 95 m a.s.l., on rock walls around cave, 10 Mar. 2015 (B.A. Huber, B. Petcharad)GoogleMaps  .

Other material examined

THAILAND: 6 ♂♂, 5 ♀♀, 2 juvs, ZFMK (5 ♂♂, 4 ♀♀, Ar 15029–30)GoogleMaps  and PSUZC (1 ♂, 1 ♀), same data as holotypeGoogleMaps  ; 1 ♂, 2 ♀♀, 3 juvs, in absolute ethanol, ZFMK ( Mal 350), same data  .

Description

Male (holotype)

MEASUREMENTS. Total body length 4.6, carapace width 1.3. Leg 1: 48.6 (11.6 + 0.6 + 11.7 + 22.8 + 1.9), tibia 2: 7.9, tibia 3: 4.5, tibia 4: 6.3; tibia 1 L/d: 95. Distance PME-PME 340 µm, diameter PME 125 µm, distance PME-ALE ~35 µm; distance AME-AME 50 µm; diameter AME 55 µm.

COLOR. Carapace pale ochre-yellow with pair of light brown marks posteriorly; ocular area and clypeus not darkened; sternum light brown with lighter marks and dark lateral margins; legs ochre-yellow with dark brown patellae and tibia-metatarsus joints; abdomen monochromous ochre-gray.

BODY. Habitus as in Fig. 53View Figs 49 – 56; ocular area slightly raised, with brushes of stronger hairs behind each PME ( Figs 68–69View Figs 68 – 78); carapace without median furrow; clypeus unmodified; sternum wider than long (0.84/0.52), unmodified. ALS with one widened, one pointed, and six smaller cylindrically shaped spigots of varying sizes ( Fig. 78View Figs 68 – 78).

CHELICERAE. As in Fig. 65View Figs 63 – 67, with pair of distal frontal apophyses provided with two to three modified (cone-shaped) hairs each ( Fig. 70View Figs 68 – 78), pair of rounded lateral processes, and pair of small indistinct proximal frontal humps.

PALPS. As in Figs 63–64View Figs 63 – 67; coxa unmodified; trochanter with large retrolatero-ventral apophysis; femur proximally widened on ventral side, with small retrolatero-dorsal apophysis; tarsus without dorsal elongation; procursus rather simple ( Figs 71–72View Figs 68 – 78), with prolateral weakly sclerotized process and transparent membranous structures; procursus tip as in Fig. 75View Figs 68 – 78; bulb with distinctive uncus and appendix with retrolateral process ( Fig. 73View Figs 68 – 78); weakly sclerotized short embolus.

LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 6%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 with> 30 pseudosegments, distally fairly distinct. Tarsus 4 comb-hairs as in Fig. 71View Figs 68 – 78.

Male (variation)

Tibia 1 in 5 other males: 10.1–12.8 (mean 11.7). Some males with white marks on abdomen.

Female

In general similar to male ( Fig. 52View Figs 49 – 56) but without stronger hairs behind PME; eye triads closer together than in male (PME-PME distance: 220 µm). Tibia 1 in 5 females: 8.7–10.1 (mean 9.3). Epigynum weakly sclerotized bulging area, only posterior area more strongly sclerotized, with small but distinct ‘knob’ ( Figs 66View Figs 63 – 67, 77View Figs 68 – 78, 99–100View Figs 90 – 101); internal genitalia as in Figs 67View Figs 63 – 67 and 101View Figs 90 – 101, with anterior arch and distinctive triangular sclerite visible through cuticle.

Natural history

This species was abundant at the type locality on vertical and slightly overhanging smooth rocks. Specimens were observed tightly pressed against the rock surface, in some cases with a small domed web nearby. When disturbed, the spiders dropped to the ground.

Distribution

Known from type locality only ( Fig. 57View Fig. 57).

ZFMK

Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"

PSUZC

PSUZC

AME

USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Pholcidae

Genus

Pholcus