Pholcus lambir Huber

Berhard A. Huber, Booppa Petchard, Charles Leh Moi Ung, Joseph K. H. Koh & Amir R. M. Ghazali, 2016, The Southeast Asian Pholcus halabala species group (Araneae, Pholcidae): new data from field observations and ultrastructure, European Journal of Taxonomy 190, pp. 1-55: 49-53

publication ID

10.5852/ejt.2016.190

publication LSID

lsid:zoobank.org:pub:urn:lsid:zoobank.org:pub:BE92596B-62D9-46CD-8486-CF6B36C640B11

persistent identifier

http://treatment.plazi.org/id/2B969B98-2995-4325-911C-D3013371FFE3

taxon LSID

lsid:zoobank.org:act:2B969B98-2995-4325-911C-D3013371FFE3

treatment provided by

Jeremy

scientific name

Pholcus lambir Huber
status

sp. nov.

Pholcus lambir Huber  , sp. nov.

urn:lsid:zoobank.org:act:2B969B98-2995-4325-911C-D3013371FFE3

Figs 195–218View Figs 193 – 199View Figs 200 – 204View Figs 205 – 210View Figs 211 – 218

Diagnosis

Easily distinguished from putatively closest known relative ( Ph. andulau  ) by very short dorsal (slightly prolateral) process on procursus ( Fig. 200View Figs 200 – 204), by much larger sclerotized teeth on male embolus ( Figs 200View Figs 200 – 204, 209–212View Figs 205 – 210View Figs 211 – 218), and by less curved anterior sclerite in internal female genitalia ( Figs 197–199View Figs 193 – 199).

Etymology

The species name is derived from the type locality; noun in apposition.

Type material

MALAYSIA-BORNEO: holotype, ♂, ZFMK ( Ar 15059), Sarawak, Lambir Hills National Park (4.198– 4.207° N, 114.034– 114.045° E), 60–150 m a.s.l., in domed webs under leaves, 22 Jul. 2014 (B.A. Huber, S.B. Huber)GoogleMaps  .

Other material examined

MALAYSIA-BORNEO: 9 ♂♂, 13 ♀♀, 1 juv., ZFMK (8 ♂♂, 12 ♀♀, Ar 15060–61)GoogleMaps  and SMK (1 ♂, 1 ♀), same data as holotypeGoogleMaps  ; 1 ♂, 2 ♀♀, in absolute ethanol, ZFMK ( Bor 200), same dataGoogleMaps  ; 3 ♂♂, 8 ♀♀, ZFMK ( Ar 15062–63), Sarawak, Niah Cave National Park, forest along main trail (3.814– 3.821° N, 113.763– 113.771° E), 20–40 m a.s.l., domed webs under leaves, 27 Jul. 2014 (B.A. Huber, S.B. Huber)GoogleMaps  .

Description

Male (holotype)

MEASUREMENTS. Total body length 5.0, carapace width 1.05. Leg 1: 48.4 (10.6 + 0.5 + 11.0 + 23.5 + 2.8), tibia 2: 6.9, tibia 3: 4.0, tibia 4: 6.2; tibia 1 L/d: 100. Distance PME-PME 380 µm, diameter PME 100 µm, distance PME-ALE ~40 µm; AME absent (only small irregular internal black marks).

COLOR. Carapace pale ochre-yellow, ocular area and clypeus brown; sternum whitish; legs ochre-yellow to orange, with brown patellae and tibia-metatarsus joints; abdomen ochre-gray with some darker marks dorsally, monochromous ventrally.

BODY. Habitus as in Fig. 195View Figs 193 – 199; ocular area slightly raised and each triad on short stalk directed obliquely dorsad ( Fig. 205View Figs 205 – 210); carapace without median furrow; clypeus unmodified; sternum wider than long (0.80/0.65), unmodified. Gonopore with four epiandrous spigots ( Fig. 216View Figs 211 – 218). ALS with one widened, one pointed, and six smaller cylindrically shaped spigots of varying sizes ( Fig. 218View Figs 211 – 218).

CHELICERAE. As in Fig. 202View Figs 200 – 204, with distinctive pair of pointed frontal apophyses directed toward each other ( Fig. 214View Figs 211 – 218); indistinct lateral processes.

PALPS. As in Figs 200–201View Figs 200 – 204; coxa unmodified; trochanter with retrolatero-ventral apophysis; femur with distinctive ventral process directed proximad; procursus relatively simple, with small semitransparent prolateral process ( Figs 207–209View Figs 205 – 210) and distinctive sclerotized and membranous distal elements; bulb large, with strong proximal sclerite, with uncus, with proximally heavily sclerotized embolus provided with strong teeth ( Figs 209–212View Figs 205 – 210View Figs 211 – 218).

LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 2%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 pseudosegments very indistinct, only distally a few visible in dissecting microscope. Tarsus 4 comb-hairs of the simplified Pholcus  - type (cf. Huber & Fleckenstein 2008), with four lateral tines ( Fig. 215View Figs 211 – 218).

Male (variation)

Tibia 1 in 9 other males: 9.6–11.7 (mean 10.5).

Female

In general similar to male ( Fig. 196View Figs 193 – 199), but eye triads on low humps and closer together ( Fig. 206View Figs 205 – 210; PME-PME distance: 200 µm), clypeus darker than in males. Tibia 1 in 20 females: 6.8–8.8 (mean 8.0). Epigynum weakly sclerotized flat plate with large unsclerotized ‘knob’ directed toward anterior ( Fig. 213View Figs 211 – 218), anterior internal sclerite visible through cuticle ( Figs 197–198View Figs 193 – 199, 203View Figs 200 – 204); internal genitalia as in Figs 199View Figs 193 – 199 and 204View Figs 200 – 204. ALS as in male ( Fig. 217View Figs 211 – 218).

Natural history

Spiders were collected from domed webs among the vegetation, at approximately 1–2 m above the ground. The apex of the sheet was connected to the underside of a leaf, but spiders hung in their webs under the leaf rather than having their bodies pressed against the leaf. In some webs, cecidomyiid flies were seen and collected in large numbers. When disturbed, the spiders vibrated vigorously. Egg-sacs were carried in front of the body ( Fig. 196View Figs 193 – 199), as in typical pholcids; they are slightly elongate and contain ~25– 30 eggs.

Distribution

Known from two localities in northeastern Sarawak ( Fig. 153View Fig. 153).

ZFMK

Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"

SMK

SMK

AME

USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Pholcidae

Genus

Pholcus