Effossana intrinseca, Domahovski & Cavichioli, 2023

Domahovski, Alexandre Cruz & Cavichioli, Rodney Ramiro, 2023, Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong & Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data, Arthropod Systematics & amp; Phylogeny 81, pp. 79-164 : 79

publication ID

https://dx.doi.org/10.3897/asp.81.e81961

publication LSID

lsid:zoobank.org:pub:5C1FDC82-8F9F-4869-ADDD-83FA96E507ED

persistent identifier

https://treatment.plazi.org/id/932FADA1-5DB6-46DA-8CC4-A68231D872C1

taxon LSID

lsid:zoobank.org:act:932FADA1-5DB6-46DA-8CC4-A68231D872C1

treatment provided by

Arthropod Systematics & Phylogeny by Pensoft

scientific name

Effossana intrinseca
status

sp. nov.

Effossana intrinseca sp. nov.

Figs 39 View Figure 39 , 40 View Figure 40 , 43C, D View Figure 43

Diagnosis.

Head, in lateral view (Fig. 39C View Figure 39 ), crown-face transition moderately thick Pygofer, in dorsal view (Fig. 39g View Figure 39 ), with acute subapical process, directed inward. Style, in lateral view (Fig. 39K View Figure 39 ), with blade very narrow near base and strongly widened towards apex; apex truncated with thin and acute process directed dorsally. Aedeagus (Fig. 39L View Figure 39 ) with shaft very long, basal portion strongly curved forming wide ring.

Measurements.

Total length: holotype male 8.6 mm; paratypes, males (n = 4) 8.6-9.2 mm, females (n = 2) 9.3-9.4 mm.

Coloration.

Head and thorax yellowish brown. Crown (Fig. 39A View Figure 39 ) with coronal suture black. Face (Fig. 39B View Figure 39 ) gena yellow, brown markings near subgenal suture, ventral and inner margins of eye. Pronotum (Fig. 39A View Figure 39 ) with brown punctures, pair of elongated and oblique black maculae behind ocelli, near anterior margin; posterior margin with black transverse band; proepimeron (Fig. 39C View Figure 39 ), with black band below lateral carina. Mesonotum (Fig. 39A View Figure 39 ) with yellow markings, pair of black ring-shaped maculae near lateral angles and pair of rounded black spots, near scutoscutellar suture. Scutellum (Fig. 39A View Figure 39 ) black with pair of large yellow spots. Forewing (Fig. 39D View Figure 39 ) with dark-brown mottling; membrane translucent light yellow; veins brown outlined by dark brown. Metatibia (Fig. 39C View Figure 39 ) with cucullate bases of setae blacks.

Description.

Head, in dorsal view (Fig. 39A View Figure 39 ), transocular width 8.1 tenths of humeral width of pronotum. Head, in frontal view (Fig. 39B View Figure 39 ), frons 1.4 × longer than wide; clypeus 1.4 × longer than wide; lateral margins parallel. Head, in lateral view (Fig. 39C View Figure 39 ), crown-face transition moderately thick, smooth medially and with few striae near of eye. Forewing (Fig. 39D View Figure 39 ) with appendix reduced, narrower than width of first apical cell. Protibia with AD row with 2-4 differentiated setae. Metatibia PD, AD and AV rows with 23-24, 12-13 and 16-18 macrosetae respectively. Metatarsomere I moderately elongated, about 3 × longer than apical width; inner row of ventral surface with 5-6 setae; pecten with 6-7 platellae. Metatarsomere II pecten with 3 platellae. Other characteristics as in generic description.

Male terminalia.

Sternite VIII (Fig. 39E View Figure 39 ) 1.4 × wider than long, lateral margins rounded; posterior margin with median third deeply excavated, bearing short median projection. Valve (Fig. 39F View Figure 39 ) 2.5 × wider than long; posterior margin narrowly excavated; in lateral view, strongly convex. Pygofer, in lateral view (Fig. 39G View Figure 39 ), 2.2 × longer than maximum height; anteroventral margin sinuous; posterodorsal and posteroventral margins slightly rounded; apex narrowed and rounded; in dorsal view (Fig. 39g View Figure 39 ), with acute subapical process, directed inward. Subgenital plate, in ventral view (Fig. 39H View Figure 39 ), elongated, 3.4 × longer than wide, maximum width near half length; inner margin straight; outer margin broadly rounded; apex rounded. Connective (Fig. 39I View Figure 39 ) with thin arms. Style, in lateral view (Fig. 39K View Figure 39 ), with blade very narrow near base and strongly widened towards apex; dorsal margin excavated; ventral margin straight, with apical third serrated; apex truncated and very wide, with thin and acute process directed dorsally. Aedeagus (Fig. 39L, M View Figure 39 ) dorsal apodeme reduced, dorsal margin excavated, lateral margins not projected laterally; apodemal process elongated and curved dorsally, dorsal margin slightly excavated medially, apex slightly broadened and rounded; shaft cylindrical and very long, basal portion strongly curved forming wide ring, portion after the ring straight; apex slightly twisted laterally, with two pair of processes directed dorsally: apical pair very short, near lateral margins and subapical pair 4 × longer than the apical pair, arising from the posterior surface. Other characteristics as in generic description.

Female terminalia.

Sternite VII (Fig. 40A View Figure 40 ) 2 × wider than long; posterolateral angles produced as far as posterior margin; posterior margin deeply excavated each side of rounded median lobe which occupies more than median third. Internal sternite VIII membranous. Pygofer (Fig. 40A, B View Figure 40 ) 1.9 × longer than maximum height; apex rounded. First valvifer (Fig. 40C View Figure 40 ) sub-rectangular 1.6 × wider than long; dorsal margin straight; posterior margin rounded. First valvula (Fig. 40C View Figure 40 ) 4 × longer than wide. Second valvula (Fig. 40E View Figure 40 ) 4 × longer than wide; dorsal protuberance rounded. Gonoplac (Fig. 40G View Figure 40 ) 3.7 × longer than wide. Other characteristics as in the diagnosis of the genus.

Material examined.

Holotype male: Brazil: Mato Grosso: “Cáceres, MT\ 27/III/1985 \ C. Ellias leg.\ Polonoroeste", "Dpt° Zool\ UF-Paraná” (DZUP) . Paratypes: 2♂, 1♀, same data as holotype ; 1♂, same data as holotype except 20.XII.1984 (DZUP); 1♀, same data as holotype except 14.I.1985 (DZUP).

Etymology.

the new species name refers to the process of pygofer directed inward.

Remarks.

Effossana intrinseca sp. nov. can be distinguish from E. circumnota sp. nov. by the pygofer (Fig. 39g View Figure 39 ) with a subapical process directed inward; the style (Fig. 39K View Figure 39 ) with ventral margin of blade without process, and the apex truncated and very wide, forming a thin and acute process directed dorsally.