Onthophagus (Onthophagus) dicranius Bates, 1887

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 182-183

Onthophagus (Onthophagus) dicranius Bates, 1887 Plate 40D

Onthophagus dicranius Bates, 1887: 72 (original description. Type locality: PANAMA, Bugaba).

Onthophagus dicranius : Gillet 1911a: 205 (complete list of species); Boucomont and Gillet 1927: 205 (catalog of species); Boucomont 1932: 298 (characters in key), 324 (distribution); Blackwelder 1944: 211 (list of species from Latin America); Howden and Young 1981: 96 (characters in key), 112 (redescription); Howden and Gill 1993: 1093 (characters in key); 1094 (redescription); Zunino and Halffter 1997: 165 (list of species); Kohlmann and Solís 2001: 164 (characters in key), 201 redescription); Medina et al. 2001: 139 (cited for Colombia); Ratcliffe 2002: 17 (cited for Panama); Morón 2003: 74 (cited for Mexico); Pulido-Herrera and Zunino 2007: 102 (catalog of species, distribution); Carvajal et al. 2011: 322-323 (cited for Ecuador); Krajcik 2012: 178 (complete list of species); Solís and Kohlmann 2012: 8 (cited for Costa Rica); Delgado and Curoe 2014: 65 (characters in key, cited for Panama).

Onthophagus (Onthophagus) dicranius : Chamorro et al. 2018: 97 (cited for Ecuador).

Type specimens.

Onthophagus dicranius Bates, 1887. The lectotype is deposited at the NHML (see Kohlmann and Solís 2001: 201). Locality: Bugabá 800-1000 feet [= 240-805 m], not examined.

Distribution.

Colombia, Costa Rica, Ecuador, and Panama.

Records examined.

LOS RIOS: CCRP [= Centro Científico Río Palenque] (5 specimens CEMT).

Literature records.

PICHINCHA [= SANTO DOMINGO DE LOS TSÁCHILAS]: Santo Domingo de los Colorados ( Howden and Gill 1993: 1093).

Temporal data.

It is unknown when this species was collected.

Remarks.

Inhabits coastal lowland evergreen forests. The collection method is unknown; however, according to Howden and Young (1981), this species was collected in Panama using rotten fruit of Gustavia sp. and human feces. There are specimens housed at the CEMT labelled as ECUADOR, Los Ríos CCRP (possibly CCRP is the accronym for Centro Científico Río Palenque; also known as Estación Científica Río Palenque).