Thyropygus resimus Attems, 1938

Pimvichai, Piyatida, Enghoff, Henrik & Panha, Somsak, 2011, A revision of the Thyropygus allevatus group. Part 3: the T. induratus subgroup (Diplopoda: Spirostreptida: Harpagophoridae), Zootaxa 2941, pp. 47-68 : 53-55

Thyropygus resimus Attems, 1938

( Figs. 3 A–E, 10A–B)

Thyropygus resimus Attems, 1938: 283

Thyropisthus resimus: Attems 1942: 80

Thyropygus allevatus resimus: Hoffman 1975: 136 , 137 figs 7–8 NOT Thyropygus confusus tenuispinus: Demange 1983 NOT Thyropisthus resimus: Demange 1961

Material examined. 2 males SYNTYPES in same vial, LAOS, Savanna Khet, leg. C. Dawydoff 1933 ( NMW 2575), 5 males, 5 females THAILAND, Kalasin Province, Sahussakhan District, Phu-Kum-Khao, 16° 41ˏ 33˝ N, 103° 31ˏ 30˝ E. 18 May 2007. S. Satan leg., ( CUMZ). 1 male THAILAND, Tak Province, Umphang District, Ta Kho Pi cave, 16° 3ˏ 14˝ N, 98° 49ˏ 16˝ E. 5 July 2009. Members of Animal Systematics Research Unit leg., ( CUMZ). 10 males, 10 females THAILAND, Loei Province, Muang District, Piya cave, 17° 32ˏ 10˝ N, 101° 42ˏ 26˝ E. 14 May 2008. R. Chanabun and members of Animal Systematics Research Unit leg., ( CUMZ). 10 males, 3 females THAILAND, Mahasarakham Province, Muang District, Ban Tha-Toom, 16° 10ˏ 43˝ N, 103° 27ˏ 4˝ E. 30 July 2010. R. Chanabun leg., ( CUMZ). 2 males, 2 females THAILAND, Kamphaengphet Province, Klonglarn District, Klong Larn waterfall National Park, 16° 7ˏ 50˝ N, 99° 16ˏ 38˝ E. 8 June 2009. S. Panha and members of Animal Systematics Research Unit leg., ( CUMZ). 5 males, 5 females THAILAND, Sakonnakorn Province, Phu-Pan District, Saerithai Cave, 17° 5ˏ 42˝ N, 103° 58ˏ 12˝ E. 10 September 2009. R. Chanabun and members of Animal Systematics Research Unit leg., ( CUMZ). 1 male, 5 females THAILAND, Phitsanulok Province, Wat Bod District, Kaeng Ched Kweaw entrance, 17° 8ˏ 32˝ N, 100° 26ˏ 28˝ E. 22 July 2008. R. Chanabun and members of Animal Systematics Research Unit leg., ( CUMZ). 1 male THAILAND, Tak Province, at Mae Usu cave, 17° 18ˏ 15˝ N, 94° 9ˏ 20˝ E. 29 May 2009, C. Sutcharit and members of Animal Systematics Research Unit leg., ( CUMZ). 1 male Laos, Vientiane Province, Houayang National Park, 18° 5ˏ N, 102° 40ˏ E. 22–24 May 2008. A. Solodovnikov and J. Pedersen leg., ( ZMUC). 6 males Laos, Champasak Province, Bolaven plateau, Muang Paxong, Ban Thongvay, 15° 14ˏ N, 106° 31ˏ E. 7–16 July 2008. A. Solodovnikov and J. Pedersen leg., ( ZMUC). 1 male Laos, Salavan Province, 20 km. East of Ban Nong Bua, 15° 40ˏ N, 106° 33ˏ E. 10–13 December 2007. S. Tarasov leg., ( ZMUC). 1 male THAILAND, Chiang Mai Province, Fang Horticult. Exp. Station. 21 October 1981. Zoological Museum Copenhagen leg., ( ZMUC). 1 male BURMA, Tenasserim, Mekane. 7–8 November 1934. R. Malaise leg., ( ZMUC).

1 male THAILAND, Loei Provine, Ban Si Than (mislabeled as Silang, Northern Thailand). 3 December 1958. B. Degerbøl leg., ( ZMUC).

Diagnosis. A species of the induratus subgroup. Differing from all other species in the subgroup by having one or more denticles in the middle of tibial spine (ti) and by having anterior coxal fold (ac) apically bent abruptly mesad with an acute narrowed, slender tip.

Description. Adult males with 57–71 podous rings, no apodous rings. Length 15–21 cm, width 8.0– 12.9 mm. Adult females with 64–71 podous rings, no apodous rings. Length 13–20 cm, width 9.4–14.1 mm. Overall color of living animal in some populations ( Fig. 10A ) dark brown, with prozonite brownish grey and epiproct reddish brown. In some populations, overall color of living animal ( Fig. 10B ) reddish brown.

Gonopods ( Figs. 3 A–E): Anterior coxal fold (ac) ( Fig. 3 A) basally slender, becoming broader towards tip, apically bent abruptly mesad, with acute, slender tip. Posterior coxal fold (pc) ( Fig. 3 B) basally with moderately high lateral paracoxites (px); mesal process (pmp) slightly lower than ac, forming shelf (sh) for accommodation of telopodite, distally directed distolaterad. Telopodite ( Figs. 3 C–E) leaving coxite over shelf of posterior coxal fold; femoral spine (fe) relatively long, curving in a horizontal plane, in situ resting along apical part of ac; tibial spine (ti) very long, slender, curving in an oblique-sagittal plane, in the middle sometimes with a single denticle (de) (rarely with several denticles) directed laterally, its tip in situ close to tip of fe and ac; apical part: lamellar lobe (ll) broad, rounded, transparent; palette (pa) simple, basally with an uncate subterminal lobe (usl) ventrally; distally with about six to eight brownish blepharochaetae (bp).

Notes. Colour: some specimens are overall dark brown, some are reddish brown; specimens from Laos have a longitudinal brown band mid-dorsally on the body, not seen in Thai specimens. The gonopods show some variation of the tip of anterior coxal fold (ac): some specimens with a relatively short, blunt tip, others with a narrower, relatively long, almost sharp tip. The femoral spine (fe) is relatively short in some specimens, but in Tak and Kamphaengphet specimens it is much longer and is distinctly crenulated along its inner curvature. The tibial spine (ti) has a denticle (de) (sometimes several denticles) on the outer margin at mid-length or slightly further distal in some specimens, others lack such denticles.

Concerning Thyropisthus resimus sensu Demange (1961) , see above under T. induratus . The specimen from “Northern Thailand Silang” recorded as T. confusus tenuispinus by Demange (1983) is in fact a T. resimus , see list of material for interpretation of ‘Silang’. A specimen from the orchid museum, Bangkok, also recorded as T. confusus tenuispinus by Demange (1983) is in fact a T. allevatus . These are so far the only records of T. confusus from Thailand, so this species should be removed from the list of Thai species.

Figures

FIGURE 3. Thyropygus resimus (specimen from Phu-Kum-Khao), gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C, D: left telopodite, posterior-mesal view. E: left telopodite, anterior-lateral view.

FIGURE 10. A, B: Thyropygus resimus, living males (specimens from A: Phu-Kum-Khao, B: Klong Larn waterfall National Park.). C, D: Thyropygus uncinatus, living male and female (specimens from C: Roi-Et (male), D: Phukhae Botanical Garden (female).