Trinomys gratiosus Moojen 1948

Wilson, Don E. & Reeder, DeeAnn, 2005, Order Rodentia - Family Echimyidae, Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press, pp. 1575-1592 : 1590

Trinomys gratiosus Moojen 1948

Trinomys gratiosus Moojen 1948 , Univ. Kansas Publ. Nus Nat. Hist., 1 (19): 379.

Type Locality: Brazil, Espirito Santo, Santa Teresa, Floresta da Caixa Dagua (19 o 55’S, 40 o 36’W) .

Vernacular Names: Gracile Atlantic Spiny-rat.

Subspecies: :

Subspecies Trinomys gratiosus subsp. gratiosus Moojen 1948

Subspecies Trinomys gratiosus subsp. bonafidei Moojen 1948

Distribution: South bank of River Doce, Espírito Santo southward to Teresopolis, Rio de Janeiro ( SE Brazil).

Discussion: Recognized as a subspecies of iheringi by Moojen (1948) and Pessôa and Reis (1996). Lara and Patton (2000) elevated gratiosus to a species, retained bonafidei as a subspecies of gratiosus , and suggested that panema was not morphologically distinct from gratiosus . Karyotype has 2n=56 and FN=108 (Pessôa and Reis, 1996).