Thomomys (Megascapheus) bottae (Eydoux and Gervais 1836)

Wilson, Don E. & Reeder, DeeAnn, 2005, Order Rodentia - Family Geomyidae, Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press, pp. 859-870 : 866

Thomomys (Megascapheus) bottae (Eydoux and Gervais 1836)

[Oryctomys] bottae Eydoux and Gervais 1836 , Mag. Zool. Paris, 6: 23.

Type Locality: USA, coast of California (restricted to vicinity of Monterey, Monterey Co., by Baird, 1857).

Vernacular Names: Botta's Pocket Gopher.

Subspecies: :

Subspecies Thomomys (Megascapheus) bottae subsp. bottae (Eydoux and Gervais 1836)

Subspecies Thomomys (Megascapheus) bottae subsp. abbotti Huey 1928

Subspecies Thomomys (Megascapheus) bottae subsp. abstrusus Hall and Davis 1935

Subspecies Thomomys (Megascapheus) bottae subsp. actuosus Kelson 1951

Subspecies Thomomys (Megascapheus) bottae subsp. albatus (Grinnell 1912)

Subspecies Thomomys (Megascapheus) bottae subsp. albicaudatus Hall 1930

Subspecies Thomomys (Megascapheus) bottae subsp. alexandrae (Goldman 1933)

Subspecies Thomomys (Megascapheus) bottae subsp. alpinus ( Merriam 1897)

Subspecies Thomomys (Megascapheus) bottae subsp. alticolus J. A. Allen 1899

Subspecies Thomomys (Megascapheus) bottae subsp. analogus Goldman 1938

Subspecies Thomomys (Megascapheus) bottae subsp. angustidens Baker 1953

Subspecies Thomomys (Megascapheus) bottae subsp. anitae J. A. Allen 1898

Subspecies Thomomys (Megascapheus) bottae subsp. aphrastus (Elliot 1903)

Subspecies Thomomys (Megascapheus) bottae subsp. aureiventris Hall 1930

Subspecies Thomomys (Megascapheus) bottae subsp. aureus (J. A. Allen 1893)

Subspecies Thomomys (Megascapheus) bottae subsp. awahnee Merriam 1908

Subspecies Thomomys (Megascapheus) bottae subsp. baileyi (Merriam 1901)

Subspecies Thomomys (Megascapheus) bottae subsp. basilicae Benson and Tillotson 1940

Subspecies Thomomys (Megascapheus) bottae subsp. birdseyei Goldman 1937

Subspecies Thomomys (Megascapheus) bottae subsp. bonnevillei Durant 1946

Subspecies Thomomys (Megascapheus) bottae subsp. borjasensis Huey 1945

Subspecies Thomomys (Megascapheus) bottae subsp. brazierhowelli Huey 1960

Subspecies Thomomys (Megascapheus) bottae subsp. brevidens Hall 1932

Subspecies Thomomys (Megascapheus) bottae subsp. cactophilus Huey 1929

Subspecies Thomomys (Megascapheus) bottae subsp. camoae Burt 1937

Subspecies Thomomys (Megascapheus) bottae subsp. canus (Bailey 1910)

Subspecies Thomomys (Megascapheus) bottae subsp. catalinae Goldman 1931

Subspecies Thomomys (Megascapheus) bottae subsp. catavinensis Huey 1931

Subspecies Thomomys (Megascapheus) bottae subsp. centralis Hall 1930

Subspecies Thomomys (Megascapheus) bottae subsp. cervinus (J. A. Allen 1895)

Subspecies Thomomys (Megascapheus) bottae subsp. chrysonotus (Grinnell 1912)

Subspecies Thomomys (Megascapheus) bottae subsp. cinereus Hall 1932

Subspecies Thomomys (Megascapheus) bottae subsp. collis Hooper 1940

Subspecies Thomomys (Megascapheus) bottae subsp. concisor Hall and Davis 1935

Subspecies Thomomys (Megascapheus) bottae subsp. confinalis Goldman 1936

Subspecies Thomomys (Megascapheus) bottae subsp. connectens Hall 1936

Subspecies Thomomys (Megascapheus) bottae subsp. contractus Durrant 1946

Subspecies Thomomys (Megascapheus) bottae subsp. convergens Nelson and Goldman 1934

Subspecies Thomomys (Megascapheus) bottae subsp. convexus Durrant 1939

Subspecies Thomomys (Megascapheus) bottae subsp. cultellus Kelson 1951

Subspecies Thomomys (Megascapheus) bottae subsp. cunicularius Huey 1945

Subspecies Thomomys (Megascapheus) bottae subsp. curtatus Hall 1932

Subspecies Thomomys (Megascapheus) bottae subsp. depressus Hall 1932

Subspecies Thomomys (Megascapheus) bottae subsp. desertorum (Merriam 1901)

Subspecies Thomomys (Megascapheus) bottae subsp. detumidus Grinnell 1935

Subspecies Thomomys (Megascapheus) bottae subsp. dissimilis Goldman 1931

Subspecies Thomomys (Megascapheus) bottae subsp. divergens Nelson and Goldman 1934

Subspecies Thomomys (Megascapheus) bottae subsp. estanciae Benson and Tillotson 1939

Subspecies Thomomys (Megascapheus) bottae subsp. fulvus (Woodhouse 1852)

Subspecies Thomomys (Megascapheus) bottae subsp. fumosus Hall 1932

Subspecies Thomomys (Megascapheus) bottae subsp. guadalupensis Goldman 1936

Subspecies Thomomys (Megascapheus) bottae subsp. homorus Huey 1949

Subspecies Thomomys (Megascapheus) bottae subsp. howelli Goldman 1936

Subspecies Thomomys (Megascapheus) bottae subsp. humilis Baker 1953

Subspecies Thomomys (Megascapheus) bottae subsp. imitabilis Goldman 1939

Subspecies Thomomys (Megascapheus) bottae subsp. incomptus Goldman 1939

Subspecies Thomomys (Megascapheus) bottae subsp. internatus Goldman 1936

Subspecies Thomomys (Megascapheus) bottae subsp. jojobae Huey 1945

Subspecies Thomomys (Megascapheus) bottae subsp. juarezensis Huey 1945

Subspecies Thomomys (Megascapheus) bottae subsp. lachuguilla Bailey 1902

Subspecies Thomomys (Megascapheus) bottae subsp. lacrymalis Hall 1932

Subspecies Thomomys (Megascapheus) bottae subsp. laticeps (Baird 1855)

Subspecies Thomomys (Megascapheus) bottae subsp. latus Hall and Davis 1935

Subspecies Thomomys (Megascapheus) bottae subsp. lenis Goldman 1942

Subspecies Thomomys (Megascapheus) bottae subsp. leucodon ( Merriam 1897)

Subspecies Thomomys (Megascapheus) bottae subsp. levidensis Goldman 1942

Subspecies Thomomys (Megascapheus) bottae subsp. limitaris Goldman 1936

Subspecies Thomomys (Megascapheus) bottae subsp. limpiae Blair 1939

Subspecies Thomomys (Megascapheus) bottae subsp. litoris Burt 1940

Subspecies Thomomys (Megascapheus) bottae subsp. lucidus Hall 1932

Subspecies Thomomys (Megascapheus) bottae subsp. lucrificus Hall and Durham 1938

Subspecies Thomomys (Megascapheus) bottae subsp. magdalenae (Nelson and Goldman 1909)

Subspecies Thomomys (Megascapheus) bottae subsp. martirensis J. A. Allen 1898

Subspecies Thomomys (Megascapheus) bottae subsp. mearnsi ( Bailey 1914)

Subspecies Thomomys (Megascapheus) bottae subsp. mewa (Merriam 1908)

Subspecies Thomomys (Megascapheus) bottae subsp. minimus Durrant 1939

Subspecies Thomomys (Megascapheus) bottae subsp. modicus Goldman 1931

Subspecies Thomomys (Megascapheus) bottae subsp. morulus Hooper 1940

Subspecies Thomomys (Megascapheus) bottae subsp. nanus Hall 1932

Subspecies Thomomys (Megascapheus) bottae subsp. navus Merriam 1901

Subspecies Thomomys (Megascapheus) bottae subsp. neglectus ( Bailey 1914)

Subspecies Thomomys (Megascapheus) bottae subsp. nesophilus Durrant 1936

Subspecies Thomomys (Megascapheus) bottae subsp. nigricans Rhoads 1895

Subspecies Thomomys (Megascapheus) bottae subsp. operarius ( Merriam 1897)

Subspecies Thomomys (Megascapheus) bottae subsp. optabilis Goldman 1936

Subspecies Thomomys (Megascapheus) bottae subsp. opulentus Goldman 1935

Subspecies Thomomys (Megascapheus) bottae subsp. osgoodi Goldman 1931

Subspecies Thomomys (Megascapheus) bottae subsp. paguatae Hooper 1940

Subspecies Thomomys (Megascapheus) bottae subsp. pascalis Merriam 1901

Subspecies Thomomys (Megascapheus) bottae subsp. pectoralis (Goldman 1936)

Subspecies Thomomys (Megascapheus) bottae subsp. peramplus Goldman 1931

Subspecies Thomomys (Megascapheus) bottae subsp. perditus Merriam 1901

Subspecies Thomomys (Megascapheus) bottae subsp. perpallidus Merriam 1886

Subspecies Thomomys (Megascapheus) bottae subsp. pervagus Merriam 1901

Subspecies Thomomys (Megascapheus) bottae subsp. pervarius Goldman 1938

Subspecies Thomomys (Megascapheus) bottae subsp. phelleoecus (Burt 1933)

Subspecies Thomomys (Megascapheus) bottae subsp. pinalensis Goldman 1938

Subspecies Thomomys (Megascapheus) bottae subsp. planirostris Burt 1931

Subspecies Thomomys (Megascapheus) bottae subsp. planorum Hooper 1940

Subspecies Thomomys (Megascapheus) bottae subsp. powelli Durrant 1955

Subspecies Thomomys (Megascapheus) bottae subsp. proximarinus Huey 1945

Subspecies Thomomys (Megascapheus) bottae subsp. pusillus Goldman 1931

Subspecies Thomomys (Megascapheus) bottae subsp. retractus Baker 1953

Subspecies Thomomys (Megascapheus) bottae subsp. rhizophagus Huey 1949

Subspecies Thomomys (Megascapheus) bottae subsp. riparius Grinnell and Hill 1936

Subspecies Thomomys (Megascapheus) bottae subsp. robustus Durrant 1946

Subspecies Thomomys (Megascapheus) bottae subsp. rubidus Youngman 1958

Subspecies Thomomys (Megascapheus) bottae subsp. ruidosae Hall 1932

Subspecies Thomomys (Megascapheus) bottae subsp. rupestris Chattin 1941

Subspecies Thomomys (Megascapheus) bottae subsp. ruricola Huey 1949

Subspecies Thomomys (Megascapheus) bottae subsp. russeolus Nelson and Goldman 1909

Subspecies Thomomys (Megascapheus) bottae subsp. saxatilis Grinnell 1934

Subspecies Thomomys (Megascapheus) bottae subsp. scotophilus Davis 1940

Subspecies Thomomys (Megascapheus) bottae subsp. sevieri Durrant 1946

Subspecies Thomomys (Megascapheus) bottae subsp. siccovallis Huey 1945

Subspecies Thomomys (Megascapheus) bottae subsp. simulus (Nelson and Goldman 1934)

Subspecies Thomomys (Megascapheus) bottae subsp. sinaloae (Merriam 1901)

Subspecies Thomomys (Megascapheus) bottae subsp. solitarius (Grinnell 1926)

Subspecies Thomomys (Megascapheus) bottae subsp. spatiosus Goldman 1938

Subspecies Thomomys (Megascapheus) bottae subsp. stansburyi Durrant 1946

Subspecies Thomomys (Megascapheus) bottae subsp. sturgisi (Goldman 1938)

Subspecies Thomomys (Megascapheus) bottae subsp. subsimilis Goldman 1933

Subspecies Thomomys (Megascapheus) bottae subsp. texensis Bailey 1902

Subspecies Thomomys (Megascapheus) bottae subsp. tivius Durrant 1937

Subspecies Thomomys (Megascapheus) bottae subsp. toltecus (J. A. Allen 1893)

Subspecies Thomomys (Megascapheus) bottae subsp. tularosae Hall 1932

Subspecies Thomomys (Megascapheus) bottae subsp. vanrosseni Huey 1934

Subspecies Thomomys (Megascapheus) bottae subsp. varus Hall and Long 1960

Subspecies Thomomys (Megascapheus) bottae subsp. vescus Hall and Davis 1935

Subspecies Thomomys (Megascapheus) bottae subsp. villai Baker 1953

Subspecies Thomomys (Megascapheus) bottae subsp. wahwahensis Durrant 1937

Subspecies Thomomys (Megascapheus) bottae subsp. winthropi Nelson and Goldman 1934

Subspecies Thomomys (Megascapheus) bottae subsp. xerophilus Huey 1945

Distribution: SW and W USA, north to Oregon, east to Colorado, and south to the Cape region of Baja California, Sinaloa and Nuevo Leon ( Mexico).

Conservation: IUCN – Lower Risk (lc).

Discussion: Subgenus Megascapheus . The taxonomic history of this, and related species has been contentious. Hall and Kelson (1959) included bottae within umbrinus but considered baileyi and townsendii as separate species, but Hall (1981) included all four of these taxa within his concept of umbrinus . Anderson (1966, 1972), Hoffmeister (1969, 1986), and Patton and co-workers ( Patton, 1973; Patton and Dingman, 1968; Patton and Smith, 1981) considered bottae (including baileyi) separate from umbrinus . Thaeler (1968 b), Patton et al. (1984), Patton and Smith (1989, 1994), and Rogers (1991 a, b) supported the specific separation of townsendii from Hall's (1981) umbrinus . As envisioned here, bottae includes a number of taxa considered by earlier workers to be full species but each of which are now relegated to subspecific status or are recognized as synonyms of other subspecies (taxa marked by asterisks in the list of synonyms, above). Hoffmeister (1969), Patton and Dingman (1968), and Patton (1973) reported on hybridization between bottae and umbrinus in S Arizona; Thaeler (1968 b), Patton et al. (1984), and Patton and Smith (1993) examined hybridization between bottae and townsendii in NE California. For subspecies, see Hall (1981), exclusive of those herein allocated to townsendii and umbrinus , and as modified for Arizona by Hoffmeister (1986) and for California by Patton and Smith (1990).