Stegosatyrus imbrialis (Weeks, 1901) Weeks, 1901

Zacca, Thamara, Mielke, Olaf H. H., Pyrcz, Tomasz W., Casagrande, Mirna M., Freitas, André V. L. & Boyer, Pierre, 2013, Stegosatyrus, a new genus of Euptychiina from the grasslands of neotropical realm (Lepidoptera: Nymphalidae: Satyrinae), Zootaxa 3682 (2), pp. 331-350 : 334-337

publication ID

https://dx.doi.org/10.11646/zootaxa.3682.2.7

publication LSID

lsid:zoobank.org:pub:AA334E13-A81E-4CD8-99F0-AFC4E87899DD

DOI

https://doi.org/10.5281/zenodo.6152149

persistent identifier

https://treatment.plazi.org/id/874187CE-027A-FFF6-A2DA-F98AFDF75519

treatment provided by

Plazi

scientific name

Stegosatyrus imbrialis (Weeks, 1901)
status

n. comb.

Stegosatyrus imbrialis (Weeks, 1901) n. comb.

( Figs 1 –4 View FIGURES 1 – 16 , 17, 20 View FIGURES 17 – 22 , 23–32 View FIGURES 23 – 32 , 84 View FIGURES 84 )

Epinephele imbrialis Weeks, 1901: 81 ; Weeks 1905: 43; Weymer 1911: 237; Gaede 1931: 236.

Euptychia biocellata Godman, 1905: 187 , 190, pl. X, fig. 8; Hayward 1958: 228; Forster 1964: 134 (as syn. of Megisto imbrialis ).

Euptychia subandina Jörgensen, 1935: 92 , pl. II, fig. 4; Forster 1964: 134 (as syn. of Megisto imbrialis ).

Megisto imbrialis (Weeks) ; Forster 1964: 134.

Pampasatyrus imbrialis (Weeks) ; Lamas 1996: 53; Lamas & Viloria 2004: 217.

Type locality. Bolivia, [province of Cochabamba], Alisuni.

Taxonomic history. Epinephele imbrialis Weeks, 1901 was described based on four syntypes from the type locality. Godman (1905) described Euptychia biocellata based on three syntypes from Tanampaya, Bolivia. In 1935, Jörgensen described Euptychia subandina without specifying the exact number of specimens examined [syntypes] collected in Catamarca (El Suncho and Esquina Grande), Argentina. Forster (1964) proposed a new combination transferring Epinephele imbrialis Weeks, 1901 to Megisto Hübner , [1819]. According to Forster (op. cit.), Euptychia biocellata Godman, 1905 and Euptychia subandina Jörgensen, 1935 are synonyms of Megisto imbrialis . Lamas (1996) transferred it to Pampasatyrus .

Diagnosis. The chromatic pattern on the dorsal face of wing of Stegosatyrus imbrialis n. comb. resembles Pampasatyrus gyrtone , being easily distinguished by HWV, by the absence of ocelli and presence of two light bands, one on the discal region between Rs- 2 A and other in the marginal region. Together with these characteristics, S. imbrialis n. comb. differs from the other species of the genus by the third segment of labial palp being longer than the first segment, and posterior portion of aedeagus is bipartite.

Redescription. MALE ( Figs 1 –2 View FIGURES 1 – 16 , 17, 20 View FIGURES 17 – 22 , 23–30, 32 View FIGURES 23 – 32 ). Head ( Figs 17, 20 View FIGURES 17 – 22 ): Antenna light brown with 34 flagellomeres; frons with a tuft of small and elongated light brown scales; labial palp with elongated black and white or light brown scales on the ventral face, third segment with a pointed apex, larger than the first segment. Thorax: with dark brown scales and a few elongated medium brown scales, prothoracic leg with femur and tibia of same length and tarsus 2 / 3 the length of tibia. Abdomen: dorso-laterally dark brown and ventrally light brown. Wings ( Figs 1 –2 View FIGURES 1 – 16 , 32 View FIGURES 23 – 32 ): dorsally dark brown; ventral face light brown. FW (length: 17–19 mm): external margin with dark brown fringes; R 1 appears on the discal cell, before dcs vein; sub-marginal region with a double black ocellus between R 5 -M 3 externally ringed with a yellow line and two white pupils; ventrally, marginal region with slight dark brown line between R 4 - 2 A, parallel to the margin, that connects with discal line surrounding the ocellus at R 4 -CuA 2 up to the discal region. HW: external margin without indentations, but with fringes similar to the FW; dorsal face with ocellus between CuA 1 -CuA 2; ventral face with a lighter band in the discal region and the other band in the marginal region between Rs- 2 A; lighter from the base of the wing base to the inner margin; sub-basal region with dark brown line between Sc+R 1 and 2 A. Genitalia ( Figs 23–30 View FIGURES 23 – 32 ): tegumen with pronounced median concavity in the anterior portion; ventral arms of tegumen straight with posterior dilatation in the median region; appendix angular reduced and curved; saccus elongated with 2 / 3 the length of gnathos; apex of valvae with little crenulation; aedeagus slightly longer than the valvae and thin, with anterior portion lanceolate and posterior portion dorsally bipartite.

FEMALE ( Figs 3 –4 View FIGURES 1 – 16 , 31 View FIGURES 23 – 32 ). Similar to male. Genitalia ( Fig. 31 View FIGURES 23 – 32 ): Papillae anales of same length as tergite VIII; sternite VIII separated from tergite VIII; lamella antevaginalis sclerotized; ductus bursae sclerotized; signa 1 / 3 the length of bursa copulatrix.

Distribution ( Fig. 84 View FIGURES 84 ). Occurs in Bolivia (Cochabamba) and Argentina (Jujuy, Salta, Tucumán and Catamarca), between 800–2,100 meters. Flies in January, February, April, May, July (probably also in March and June).

Host plant. Poaceae ( Hayward 1960, 1969).

Examined material. BOLIVIA, Cochabamba: Via Cochabamba (El Fuerte), 2,000m, 13, 27.VIII. 2000, Wojtusiak leg. (DZ 21.563); Santa Cruz: Samaipata (El Fuerte), 2,000m, 73 and 2 Ƥ, 27.VIII. 2000, T. Pyrcz & J. Wojtusiak leg. ( MZUJ, DZUP 21.423); ARGENTINA, Catamarca: Singuil, 2,100m, 33, without date, without collector ( MLPA); Tucumán: 13, V. 1947, Breyer leg. (DZ 21.332), San Javier, 1,200m, 1 without abdomen 20.I. 1970, Mielke leg. (DZ 21.563), 53, without date, without collector ( MLPA 025/ 1178, MLPA 025/ 1180, MLPA 025/ 1183, MLPA 025/ 1190, MLPA 025/ 1192), Racó 43, without date, without collector ( MLPA 025/ 1179, MLPA 025/ 1184, MLPA 025/ 1185, MLPA 025/ 1187), Villa Nougues 13 21.I. 1931, without collector ( MLPA 025/ 1194), Salta: 3 without abdomen, 14.IV. 1969, Heimlich leg. (DZ 24.912, DZ 21.570, DZ 21.577), 800m, 13, 5.VI. 1933, without collector ( MLPA 025/ 1176), Valderramas 23, XII. 1935, without collector ( MLPA 025/ 1177, MLPA 025/ 1181), Jujuy: Zapla, 13, without date, without collector ( MLPA 025/ 1186), 13, 10.1939, without collector ( MLPA 025/ 1188).

Male lectotype: E. imbrialis A. G. Weeks Jr , Bolívia / Alezuni, Bolívia, Aug. 8, 1899, Coll. A. G. Weeks, Jr./ type / 160 / 7758 / Lectotypus / Epinephele imbrialis Weeks, 1901 , Zacca, Mielke & Pyrcz det. 2013.

Remarks. The lectotype designation was based on the photograph of the male syntype of Epinephele imbrialis deposited in The British Museum of National History ( BMNH), available on the website “Butterflies of America ” (http://butterfliesofamerica.com/L/ Nymphalidae .htm). The remaining specimens (syntypes) are designated paralectotypes. This will prevent any future confusion regarding the identity of this taxon, and will thus maintain taxonomic stability as recommended by the International Commission on Zoological Nomenclature (1999: 83) in their Code (Recommendation 74 A).

DZUP

Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Lepidoptera

Family

Nymphalidae

Genus

Stegosatyrus

Loc

Stegosatyrus imbrialis (Weeks, 1901)

Zacca, Thamara, Mielke, Olaf H. H., Pyrcz, Tomasz W., Casagrande, Mirna M., Freitas, André V. L. & Boyer, Pierre 2013
2013
Loc

Pampasatyrus imbrialis

Lamas 2004: 217
Lamas 1996: 53
1996
Loc

Euptychia subandina Jörgensen, 1935: 92

Forster 1964: 134
1964
Loc

Megisto imbrialis

Forster 1964: 134
1964
Loc

Epinephele imbrialis

Gaede 1931: 236
Weymer 1911: 237
1911
Loc

Euptychia biocellata

Forster 1964: 134
Hayward 1958: 228
Godman 1905: 187
1905