Helosciomyza fuscinevris

McAlpine, David K., 2012, Notes and Descriptions of Australian Helosciomyzidae or Comb-winged flies (Diptera: Schizophora), Records of the Australian Museum 64 (1), pp. 51-70 : 59

Key to Australian species of fuscinevris group

1 Pteropleuron (anepimeron) setulose; hind tibia without dark zone near base; male: abdominal pleural membrane with group of minute setulae near spiracle 1 (often absent in female); surstyli as in Figs 16, 17 (slightly variable) ....................................................................................... fuscinevris , p. 60

—— Pteropleuron bare; hind tibia usually with brown to black sub-basal or basal zone; no setulae present near abdominal spiracle 1 (male, female); male surstylus variable in shape ...................................................................................... 2

2 Hypopleuron (metepisternum) with c. 2–12 small black setulae; abdominal sternite 1 setulose; antennal segment 3 tawny-orange, without or with very indistinct darker zone; male: surstylus as in Fig. 15 ......................................................................................................................... bickeli , p. 60

—— Hypopleuron and sternite 1 without setulae; antennal segment 3 often with darker brown to blackish zone; male: surstylus variable ................................................ 3

3 Fore femur entirely pale tawny-yellow on basal half or more (significantly including the sometimes concealed anterior surface) ............................................... 4

—— Fore femur largely tawny-yellow, anterior surface with distinct brown zone extending to near base (also brown distally) .............................................................. 6

4 Antenna 3 brown to blackish on entire distal half or more; male: hind basitarsus with ventrobasal dense, non-linear cluster of short, thick, black suberect spinules; surstylus as in Fig. 22 ........................................... australica , p. 64

—— Antenna 3 usually tawny orange at and below apex, more brownish dorsally between apex and aristal insertion and on outer lateral surface; male: hind basitarsus without such cluster of short, black spinules, occasional black setulae in this position sparse and distally inclined; surstylus otherwise ...................................................................................... 5

5 Fore femur with conspicuous dark brown subapical spot on posterior surface; mesoscutum with pruinescent dorsocentral stripe not covering and emphasized by a darker cuticular dorsocentral stripe (pruinescent stripe faded in some preserved specimens); male: surstylus as in Fig. 21 ............................................................................................... neboissi , p. 63

—— Fore femur with only indistinct indication of dark subapical posterior spot (dark subapical anterior spot well developed); grey-pruinescent dorsocentral and other such longitudinal stripes of mesoscutum with underlying dark brown cuticular stripes, which tend to remain when pruinescent stripes are faded; male: surstylus as in Fig. 18 .............................................................................................. subacuta , p. 63

6 Antenna 3 tawny-orange, with only limited dorsal reddish brown zone; male: surstylus as in Fig. 23 ........................................................................... steyskali , p. 64

—— Antenna 3 dark brown to blackish on distal half or more, or at least on a substantial dorsal zone (sometimes less, as in H. driesseni ); male: surstylus otherwise ............................................................................................................... 7

7 Occipital region of head with broad supracervical zone of blackish cuticle (seen through paler pruinescence) usually with, on each side, a dorsolateral extension partly bounded medially

Figures

Figures 14–18. (14) Neosciomyza peckorum (Boyd River), terminal part of epandrium and associated structures, anterior view, right anterior epandrial process cut away for clarity, scale = 0.5 mm. (15) Helosciomyza bickeli (Cobcroft Creek), left surstylus, outer view, scale = 0.2 mm. (16) H. fuscinevris (Clarence, near Lithgow), left surstylus, outer view. (17) same specimen, right surstylus, outer view. (18) H. subacuta (Wannon River), left surstylus, outer view. Figs 15–18 all same scale. aep, anterior epandrial process; bsp, basal surstylar process; c, cercus; ep, epandrium; ss, surstylus.

Figures 19–26.(19) Helosciomyza steyskali, protandrogram.(20) H.australica, epandrium and associated structures, posterior view, semi diagrammatic, setulae omitted. (21) H. neboissi (Pelion), left surstylus, outer view, scale = 0.2 mm. (22). H. australica (Picton Lakes), left surstylus, outer view. (23). H. steyskali (Boyd River), left surstylus, outer view. (24). H. obliqua (Mount Donna Buang), left surstylus, outer view. (25). H. driesseni (holotype), left surstylus, outer view. (26). H. macalpinei (holotype), left surstylus, outer view. Figs 21–26 all to same scale. c, cercus; ep, epandrium; pbf, posterobasal foot of surstylus; ss, surstylus; s5–s8, sternites 5 to 8; t5, t6, tergites 5 and 6.