Sisyromyia elongata, Li & Yeates, 2019

Li, Xuankun & Yeates, David K., 2019, Revision of the Australian bee fly genus Sisyromyia White, 1916 (Bombyliidae, Bombyliinae, Acrophthalmydini), Zootaxa 4711 (2), pp. 201-244 : 216-219

5. Sisyromyia elongata sp. nov.

( Figs 10 , 11 )

Type specimens. Holotype ♂ Australia, WA, S31°35’ E115°48’, Cypress R [oa]d, Pinjar, 31 May 1981, MJ Smart . Paratypes ♂ (3), Australia, WA, same data as holotype .

Other specimen examined. Australia, WA, ♂ (1) ♀ (1), S31°49’ E115°57’, Henley Brook , 16 May 1981, MJ Smart . ♀ S31°38’ E115°51’, Dasypogon R [oa]d, Pinjar, 16 May 1981, MJ Smart

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Diagnosis. Medium-sized fly, body with mostly brownish hairs. Wing membrane slightly infuscated and darker on anterior half. Scutellum brownish yellow. Integumental colour of abdominal tergites black except posterolaterally brown. Outer apex of gonocoxite long.

Description. Male. Body length 9.6–10.8 mm, wing length 9.1–10.1 mm.

Head. Head about 2.2 x wider than long, mostly blackish with thick pale pruinescence and covered in admixed white to golden scales and black hairs. Eyes dichoptic, narrowly separated width of ocellus. Frons short, 1.5 x length of ocellar tubercle, upper narrow and black; lower half triangular, 2.0 x length of upper half, with thick pale pruinescence and black hairs admixed with golden scales. Ocellar tubercle slightly raised, blackish brown to black with grey pruinescence, with long black hairs. Face with thick pale pruinescence and long black hairs admixed with some golden to white scales, parafacial area with more white scales. Gena with thick pale pruinescence and long white scales. Clypeus with thick pale pruinescence and otherwise bare. Occiput with thick pale pruinescence, except dorsal area with sparse pruinescence, with black hairs admixed with some golden scales behind ocellar tubercle. Posterior eye margin slightly convex. Antennal scape and pedicel black with thick pale pruinescence, scape with long black hairs, pedicel with short black hairs; flagellum black without pruinescence, subapex with 3–5 long hairs. Scape 3.2 x as long as wide, and 2.7 x as long as pedicel, uniform from base to apex. Pedicel 1.3 x as long as wide. Flagellum 12.0 x as long as wide, 1.8 x as long as scape + pedicel, 2.6 x as long as scape, conical and slightly laterally compressed, one-segmented with apical stylus ( Fig. 10c ). Palpus long, just extending beyond oral cavity, brown with yellow hairs, one-segmented, without palpal pit. Mouthparts slender, 4.5 x as long as eye length, 2.9 x as long as head length, labellum thin and filiform ( Fig. 10h ).

Thorax. Integumental colour of scutum mostly black with sparse pale pruinescence except postalar callus brown, dorsocentral brown pruinescence stripe present. Scutum covered with short brown hairs admixed with few short black hairs, lateral with some white scales; hairs denser on anterior half. Four brown notopleural setae present. Scutellum brownish yellow with sparse pale pruinescence, with short brown hairs admixed with some short black hairs, and with white hairs laterally, black hairs denser and longer posteriorly. Pleura black with thick pale pruinescence, mostly covered in long pale yellow hairs, except dorsal half of anepisternum with brown hairs admixed with some black hairs, anepimeron, meron, laterotergite and mediotergite bare.

Legs. Legs brown. Femora cover in white to yellow scales, and with long black hairs admixed with some white hairs. Fore femur with two anteroventral bristles on apical half; mind and hind femora with one row of anteroventral bristles and some separate bristles on apical half. Bristles and other hairs on legs brown. Fore tibia 1.7 x longer than fore basitarsus, mid tibia 2.0 x longer than mid basitarsus, hind tibia 2.0 x longer than hind basitarsus.

Wings. Wing membrane infuscated on anterior margin, with darker towards wing base. Cell r 5 open, M 1 close to R 5; cell br nearly as long as cell bm, crossvein r-m arising from base of cell dm; crossvein m-m nearly as long as crossvein r-m; cell cup open ( Fig. 10d ). Haltere stem and knob dark yellow.

Abdomen. Integumental colour of tergites black except posterolaterally brown, mostly with sparse pale pruinescence, but tergites 1–2 with thick pale pruinescence. Tergite 1 with dense white to pale yellow hairs, admixed with some black hairs laterally; tergite 2 with long pale yellow hairs on anterior half, and with long brown hairs admixed with long black hairs on posterior half and laterally; tergites 3–7 with long brown hairs and long black hairs, admixed with few golden scales; tergites 4–7 with median stripe consisting of dense, decumbent short white scales. Sternites black except posterior half yellow, with sparse pale pruinescence, covered with golden scales admixed with black hairs. Genitalia. Epandrium anterior and posterior margins slightly concave ( Fig. 11d ). Lateral margin of gonocoxite nearly straight ( Figs 11a,b ); lateral ejaculatory process wide; inner apex of gonocoxite short and rounded, outer apex of gonocoxite elongate and narrow; gap between inner apices of gonocoxites medium; phallus nearly as long as gonocoxite ( Fig. 11c ).

Female. Body length 7.7–10.0 mm, wing length 10.0– 12.8 mm. Very similar to male, except frons black with thick pale pruinescence, 3.0 x as wide as ocellar tubercle, frons with golden scales admixed with long blackish brown hairs ( Fig. 10l ). Around 30 acanthophorite spines present on each side of tergite 9+10 ( Fig. 11g ).

Remarks. Sisyromyia elongata sp. nov. is similar to Sisyromyia albisquama sp. nov., but differs in having the body hairs mostly brownish; scutellum brownish yellow; anterior margin of scutellum without a white band; abdominal tergites 3–7 without white scales laterally; and outer apex of gonocoxite long.

Etymology. The specific name refers to the outer apex of gonocoxite elongate.

Distribution. Australia (WA).

Figures

FIGURE 10. Sisyromyia elongata sp. nov. External (a–i (♂); j–o (♀)): (a, j) dorsal; (b, k) lateral; (c) antenna, lateral; (d) wing; (e, l) head, dorsal; (f, m) head, frontal; (g, n) head, profile; (h, o) head, lateral; (i) abdomen, dorsal. Scale bars = 1 mm (a, b, d–o); = 0.1 mm (c).

FIGURE 11. Sisyromyia elongata sp. nov. Genitalia (a–e (♂); f, g (♀)): (a) genitalic capsule, dorsal; (b) genitalic capsule, ventral; (c) genitalic capsule, lateral; (d) epandrium, dorsal; (e) epandrium, lateral; (f) sternite 8, ventral; (g) genitalia and spermathecae. Scale bars = 0.1 mm.