Tokyosoma bellum, Mikhaljova, Elena V. & Lim, Kil-Young, 2008

Mikhaljova, Elena V. & Lim, Kil-Young, 2008, The millipede family Diplomaragnidae Attems 1907 in the Korean Peninsula, with the descriptions of a new genus and new species (Diplopoda, Chordeumatida), Zootaxa 1925, pp. 51-61 : 55-56

Tokyosoma bellum sp. n.

Figs 7–9

Material examined: Holotype: 1 male (ChNU), Yangsan, Gyeongsangnam–do, South Korea, 28 December 1990, leg. K.Y. Lim; Paratypes: 1 male, 2 females (ChNU), 1 male, 1 female ( IBSS), 1 male, 1 female ( ZMUM), same locality as for holotype, 28 December 1990, leg. K.Y. Lim. One male– paratype is devoid of head. Long pointed outgrowth of the fused mesal sheath processes of posterior gonopod colpocoxites of one from dissected gonopods is broken.

Diagnosis: Differs from congeners mainly by shape (like cup with long pointed outgrowth) of mesal sheath structure of posterior gonopods, very low, wave–shaped lateral sheath processes of posterior gonopod colpocoxites and scarcely subbasally fused colpocoxites.

Description: Male. Length 7.0–7.5 mm, width about 0.7 mm. Coloration in alcohol pale with feebly marked beautiful design of light brown spots and strips on metazona and head. Legs with marbled brown distal part. Eyes black. Antennae brown.

Body with 29 segments. Head covered with long and short setae, vertigial suture visible. Eye patches subtriangular, each composed of 14–16 ocelli. Collum semicircular. Body width gradually increasing until somite 7, body parallel–sided on somites 8–21(22), gradually tapering posteriorly. Beginning from somite 2(3) somites with normally developed paraterga which gradually grow less distinct toward hind part of body. Metazonital macrochaetae in transverse row on somites 26–28, like elongate triangle on preceding somites. Macrochaetae long, pointed apically.

Legs long and slender. Each claw of legs 1–2 at base with two small additional claws dorsally and long setiform outgrowth ventrally. Leg pairs 3–7 not larger than other walking legs. Leg pairs 3–7 with small group of funnel–shaped tarsal papillae apically near claw. Each claw of leg pairs 3–7 with long setiform outgrowth ventrally only i.e. without dorsal additional small claws. Leg pair 10 with several tarsal papillae subapically. Leg pair 11 with a few or without tarsal papillae. Midbody legs without tarsal papillae. Each claw of postgonopodal legs (including leg pairs 10 and 11) at base with long setiform outgrowth ventrally and two small additional claws dorsally; this additional claws gradually missing toward very hind legs. Each claw of very hind legs with long setiform outgrowth ventrally only. Legs 10 and 11 with coxal glands. Coxa 10 with ventral fovea of which caudomesal edge extended into thin process ( Fig. 7 ). Coxa 11 without modifications (excluding opening of coxal gland ventrally). Trochanter 11 with small conical process directed caudad.

Anterior gonopod telopodite 1–segmented, flagelliform, its distal part positioned inside elevated sheaths and somewhat protruding outside, apex curved, pointed, with tiny tooth subapically ( Fig. 8 ). Posterior gonopod colpocoxites scarcely fused subbasally. Each colpocoxite divided into broad median and narrow lateral branches; last one some flattened on sides. Mesal sheath processes of posterior gonopod colpocoxites fused medially into single cup–shaped structure (ms) with long pointed outgrowth directed caudoventrally. Lateral sheath processes of colpocoxites (lp) very low, virtually undeveloped, wave–shaped. Apex of colpocoxite curved posterad. Posterior gonopod angiocoxite with pear–shaped globule in posterior view, depressed centrally in anterior view. Posterior angiocoxal process absent. Angiocoxite in anterior view with one process; anterior angiocoxal process long, its distal portion sheathed by colpocoxite on anterior side ( Fig. 9 ). Posterior gonopod telopodite with short femur.

Female. Length 8–8.5mm, width 0.8–0.9 mm. Body with 29 segments. Ocelli 16–18. Vulvae not dissected.

Etymology: The specific epithet refers to the beautiful design of light brown spots and strips on metazona and head.

Figures

FIGURES 7 – 9. Tokyosoma bellum sp. n., male. 7, coxa 10; 8, gonopods, caudal view; 9, gonopods, front view; ms, mesal sheath processes of posterior gonopod colpocoxites; lp, lateral sheath process of posterior gonopod colpocoxite. Scale in mm.