Ummidia pustulosa (Becker, 1879)

Godwin, Rebecca L. & Bond, Jason E., 2021, Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae), ZooKeys 1027, pp. 1-165 : 1

Ummidia pustulosa (Becker, 1879) Fig. 39 , Map 4

Pachylomerus pustulosus Becker, 1879: 140.

Pachylomerus pustulosus Becker, 1881: 44.

Type material.

Female holotype presumed lost. NEOTYPE 1♀ designated here (NHMUK1071) from Amula (now Jalisco), Mexico, 20.6934 -103.356 7, 1550 m a.s.l., H.H. Smith 1898, NHMUK. The original type locality is simply listed as "Guanajuato, Mexico" in the original description which neighbors the locality of the neotype designated, Jalisco, Mexico. Given the carapace length to width ratio noted in Becker’s (1879) description and the illustration of the sternum ( Becker 1881, fig. 2b) we can be reasonably confident that designation of this specimen as the neotype is consistent with what is known of the originally described specimen.

Diagnosis.

Females of U. pustulosa can be differentiated from U. pesiou and U. rodeo by having the carapace and sternum 1.2 × long as wide (roughly long as wide in other species) and from U. gabrieli by having relatively more endite cuspules (43 vs 28).

Female neotype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen. General coloration. Carapace, chelicerae, and legs dark reddish brown 2.5YR 3/4. Abdomen brown 7.5YR 4/3, spinnerets strong brown 7.5YR 4/6. Cephalothorax. Carapace 9.44 long, 7.99 wide. Pars cephalica 6.36 long. Foveal groove slightly procurved, 0.21 long, 1.7 wide. Eye tubercle low. AER procurved. PER recurved. Eye group 0.93 long, 1.59 wide, AME 0.34, PME 0.19, ALE 0.49, PLE 0.32. Sternum moderately setose, STRl 5.92, STRw 4.85. Chelicerae with anterior row comprising seven teeth, posterior margin with seven teeth. Palpal endites with 20 cuspules spread across proximal half of endite and 23 smaller cuspules distally, ENDw 2.08, ENDl 3.5. Labium with four cuspules, LBw 2.04, LBl 1.48. Rastellum with very many strong spines on process and along margin continuing up cheliceral face approx. length of rastellar process. Abdomen. Evenly setose. Legs. F1 5.66; F1w 1.8; P1 3.52; Ti1 3.3; Mt1 2.46; Tr1 1.43; F3 4.93; F3w 2.57; P3 3.16; Ti3 2.91, Sd3 1.84; Mt3 2.06; Tr3 1.98; F4 5.75; F4w 2.66; P4 3.35; Ti4 3.33; Mt4 3.31; Tr4 1.82. Retrolateral face tarsus IV with very defined comb over length of tarsus. Pedipalps. PF 5.32, PP 2.99, PTi 3.29, PTr 3.1. Spermathecae curved laterally with distal coil, bulbs facing laterally.

Variation, females

(n = 3). CL 8.65-9.53, 9.21 ± 0.28; CW 7.5-8.62, 8.04 ± 0.32; Cap 5.63-6.36, 6.05 ± 0.22; ENDl 0.93-1, 0.96 ± 0.02; ENDw 1.59-1.84, 1.68 ± 0.08; STRl 5.27-5.92, 5.65 ± 0.19; STRw 4.31-4.85, 4.64 ± 0.17; LBl 1.48-1.65, 1.56 ± 0.05; LBw 1.62-2.04, 1.87 ± 0.13; F1 3.38-5.66, 4.63 ± 0.67; F1w 1.89-2.06, 1.97 ± 0.05; P1 3.03-3.58, 3.38 ± 0.18; Ti1 2.96-3.33, 3.2 ± 0.12; Mt1 2.17-2.59, 2.4 ± 0.12; Tr1 1.28-1.43, 1.34 ± 0.05; F3 4.39-4.93, 4.71 ± 0.16; F3w 2.31-2.65, 2.51 ± 0.1; P3 2.9-3.5, 3.19 ± 0.17; Ti3 2.78-2.97, 2.89 ± 0.06; Mt3 1.96-2.06, 2.03 ± 0.03; Tr3 1.88-1.99, 1.95 ± 0.03; F4 5.59-6.48, 5.94 ± 0.28; F4w 2.33-2.75, 2.58 ± 0.13; P4 3-3.55, 3.3 ± 0.16; Ti4 3.33-3.57, 3.45 ± 0.07; Mt4 3.16-3.47, 3.31 ± 0.09; Tr4 1.57-1.93, 1.77 ± 0.11; PF 4.33-5.32, 4.98 ± 0.32; PP 2.72-3.17, 2.96 ± 0.13; PTi 2.88-3.51, 3.23 ± 0.18; PTr 2.49-3.1, 2.82 ± 0.18.

Males.

Unknown.

Material examined.

Mexico: Jalisco: Amula (now Jalisco), Mexico, 20.6934 -103.356 7, 1550 m a.s.l. (NHMUK1071, 1898, 1♀, HH Smith, NHMUK); Chamela Biological Station , 19.5307 -105.0828 6, 13 m a.s.l. (UMM0465, 21.vii.1989, 1♀, JG Rozen, AMNH) ; Sinaloa: 35 mi N of Los Mochis , former desert floor within 0.5 mi of the ocean, 26.2772 -109.194 76, 15 m a.s.l. (UMM0222, iii.1961, 1♀, WW Gibson, MCZ) .

Figures

Map 4. Locality records for western United States and northern Mexico species.

Figure 39. Ummidia pustulosa (Becker, 1879) from Sonora, Mexico. Female neotype (BMNH 1071) A female habitus illustration B cleared spermathecae. Scale bars: 4.0 mm (A), 1.0 mm (B).