Arianops, Brendel from the, 1893

Carlton, Christopher E., 2008, Eight New Species ofArianopsBrendel from the Southeastern United States with an Updated Key and Notes on Additional Species (Coleoptera: Staphylinidae: Pselaphinae), The Coleopterists Bulletin 62 (2), pp. 297-323 : 297-323

Key to Species of the Genus Arianops

The following is based on Barr’s key that was included in his revision of the genus, with the addition of ten subsequently described species and other slight modifications ( Barr 1974). Many of the characters given are limited to the male sex. Females of species for which males are unknown are included in the key

insofar as possible, but the geographic origin of such specimens should be considered in identifying them.

1. Abdomen convex, $ 0.75 X as thick as wide (e.g., Figs. 3–4 ); often cavernicolous species from southwestern Virginia, Tennessee and Alabama, and larger, convex species from Appalachian Mountains, Ouachita Mountains , or east Texas ....................................... 2

1 9. Abdomen flattened, # 0.65 X as thick as wide (e.g., Figs. 5–6 ); more flattened species known from southwestern North Carolina, northeastern Georgia, and southwestern Alabama (henroti species group)...... 8

2(1). Head without lateral vertexal carinae ( Fig. 30 ), or carinae attenuated well before vertexal foveae ( Fig. 24 ), vertexal foveae small and shallow..... 3

2 9. Head with distinct lateral vertexal carinae extending nearly to vertexal foveae; vertexal foveae normally developed ( Fig. 17 )................... 6

3(2). Pronotum longer than wide............................................... 4

3 9. Pronotum length and width subequal.................................... 5

4(3). Pronotum with prominent median basal fovea; length 3.1 to 3.6 mm; larger, darker species inhabiting higher mountains of southwestern North Carolina (alticola species group)................................. 15

4 9. Pronotum usually without median basal fovea, rarely with small, shallow foveoid depression; length 2.3 to 3.0 mm; smaller, paler species inhabiting caves and ravines of southwest Virginia to central Tennessee and Alabama (cavernensis group, major part).......................... 16

5(3). Size large, 3.6 to 4.2 mm, robust; aedeagus about 0.6 mm long (gigantea species group).............................................. A. gigantea Barr

5 9. Smaller species, 2.9 to 3.2 mm, more slender; aedeagus about 0.4 mm long (neglecta species species group)..................................... 27

6(2). Pronotum with median basal foveae...................................... 7

6 9. Pronotum without median basal foveae; Jackson County, Alabama (cavernensis group, in part)........................... A. barri new species

7(6). Pronotum longer than wide, median fovea flanked on either side by elevated, sometimes feebly carinate ridge; paler, more slender species, males with small spine on third or fourth ventrite or spine absent, genital depression not heavily sclerotized and blackened; Great Smoky, Unicoi, and Nantahala Mountains of North Carolina and Tennessee (nantahala group)...................................................................... 31

7 9. Pronotum as long as wide, median fovea flanked on either side by prominent tubercle, usually with small spine; darker, more robust species, male with large spine or shelf on third ventrite; western Pennsylvania and Pisgah Ledge, Ouachita Mountains of Arkansas and Oklahoma, or east Texas (amplyoponica species group)............... 35

8(1). Pronotum with small but distinct median fovea near base............. 9

8 9. Pronotum without median fovea near base............................. 11

9(8). Head with lateral vertexal carinae short or interrupted................ 10

9 9. Head with long, conspicuous, lateral vertexal carinae continuing behind level of vertexal foveae; Rabun, Towns, Union, and White Counties, Georgia...................................................... A. obliqua Barr

10(9). Head with lateral vertexal carinae short; occipital carina moderately long; no median longitudinal ridge behind median fovea of pronotum; Cheoah Bald area , Graham County, North Carolina ... A. norithe Barr

Figures

Figs. 1–4. Dorsal and lateral views, Arianops new species holotypes. 1–2) A. obrieni; 3–

Figs. 5–8. Dorsal and lateral views, Arianops new species holotypes. 5–6) A. clintoni; 7–

Figs. 28–33. Arianops new species. 28) A. folkertsi, head; 29) A. folkertsi, ventrites 3–4; 30) A. gorei, head; 31) A. gorei, ventrites 3–4; 32) A. hiltenae, head; 33) A. hiltenae, ventrites 3–4.

Figs. 17–27. Arianops new species. 17) A. obrieni, head; 18) A. obrieni, female ventrites 5–6; 19) A. carteri, head; 20) A. carteri, mesothoracic coxa-femur; 21) A. clintoni, head; 22) A. clintoni, mesothoracic coxa-femur; 23) A. clintoni, ventrites 3–4; 24) A. ashei, head; 25) A. ashei, ventrites 3–4; 26) A. barri, head; 27) A. barri, ventrites 3–4.