Amphineurus (Rhamphoneurus) falcatus, Santos & Santos & Ribeiro, 2022

Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha, 2022, Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae), Arthropod Systematics & amp; Phylogeny 80, pp. 439-494 : 439

publication ID

publication LSID

persistent identifier

taxon LSID

treatment provided by

Arthropod Systematics & Phylogeny by Pensoft

scientific name

Amphineurus (Rhamphoneurus) falcatus

sp. nov.

3.3.11. Amphineurus (Rhamphoneurus) falcatus sp. nov.

Material examined.

Holotype: ♂, Chile, Osorno , Bahia Mansa [40°35′S 73°44′W], South of Pucotrihue, 12-IV-1968, Peña (USNM) GoogleMaps *. Paratypes: 1 [sex unkown]: Chile, Chiloé Is., Aulen [42°2′S 74°1′W], 8-II-1952, L.P. Guzman (USNM); 1 ♂, Chile, Chiloé Is., Aulen [42°2′S 74°1′W], 8-II-1952, Peña (USNM)*; 1 [sex unkown]: Chile, Chiloé Is., Aulen , Duhatao [41°58′S 74°02′W], 6/ 8-II-1952, Peña (USNM) GoogleMaps *.


The word Amphineurus falcatus is Latin for "armed with scythes". This name is in reference to the sheath of aedeagus, which resembles a scythe.


This species is distinguished by absence of markings near R2 and R2+3+4 longer than R2+3. Furthermore, this species has distinctive male terminalia with a curved sheath of aedeagus, only the right mesal lobe folded, and the longest lobule of the subgenus, as long as the remainder of the lobe of gonostylus.


Wing length 6.08 mm, width 1.86 mm. - Coloration: General coloration blackish brown. Thorax blackish brown, pleura gray. Scutum without stripes. Halter ochreous. Coxae brownish yellow and legs dark brown. - Head (dorsal view Fig. 17B View Figure 17 , ventral view Fig. 17A View Figure 17 ): Rostrum bifurcated with short appendices; first palpal segment as long as terminal segment; scape as thick as pedicel; last flagellomere as long as penultimate. - Thorax (lateral view Fig. 17E View Figure 17 , dorsal view Fig. 17F View Figure 17 ): Division line of laterotergite sinuous and almost vertical; anatergite as tall as katatergite. Halter with long dilated knob; length of knob half-length of stem of halter. Wing (Fig. 17D View Figure 17 ) without distinctive markings; veins near fork of bM (Fig. 17C View Figure 17 ) visible; R2 faded; r-m shorter than M1+2 and basal deflection of M1; M1+2 shorter than basal deflection of M1; R2+3 at least 2.5 × longer than basal deflection of R5; R2+3+4 longer than R2+3. - Male terminalia (Fig. 17G View Figure 17 ): Male tergite IX darkened along posterior margin, with wide V-shaped median notch; posterolateral lobes nearly triangular, obtuse at apex. Ventral branch of gonocoxite expanded on distal tip. Dorsal branch of gonocoxite shorter than half-length of ventral branch. Lobe of gonostylus with rounded lobule; lobule longer than wide, longer than half-length of lobe of gonostylus and wider than stem. Clasper of gonostylus with similarly shaped branches: lateral branch rounded; medial branch oblong. Mesal lobes of gonocoxite asymmetrical; left mesal lobe straightened and pointed; right mesal lobe bent at right angle. Sheath of aedeagus curved, darkened for most of length.


Specimens of this new species were previously identified by C.P. Alexander as A. (R.) nullus Alexander. This species resembles A. (R.) nullus but differs mainly in the attachment of m-cu, shape of male tergite IX, prescutum and left mesal lobe.