Trichoceridae Rondani, 1841

Shcherbakov, Dmitry E. & Azar, Dany, 2019, A new subfamily of Trichoceridae (Diptera: Tipulomorpha) from Early Cretaceous Lebanese amber, Zootaxa 4563 (2), pp. 311-324 : 312

Family Trichoceridae Rondani, 1841

Revised diagnosis. Tipulomorpha with distinct basal R and M bends opposed at atrophied MA (=arculus), 1A (2A of traditional system) shortened (its distal portion often subtransverse), ocelli developed, scutoscutellar suture obscure, and female cerci (when elongate) curved downwards. Pedicel somewhat enlarged, larger than scape, usually contrasting to setaceous flagellum (except for some Jurassic genera with moniliform antennae).

Composition. Trichocerinae Rondani, 1841 , Paracladurinae Krzemińska, 1992 , Kovalevinae Krzemińska, Krzemiński et Dahl, 2009 and Ewauristinae subfam. nov.

Remarks. The following characters, not found in the new genus, could no more be considered as diagnostic for Trichoceridae : crossvein sc-r in proximal position, R with 3 long branches, and R2 crossvein-like near the R3 base. In fact, R2 is not proximal also in several other trichocerid taxa: it is equidistant between R2+3 origin and R1 apex in a few species, or even slightly nearer to the R1 apex in modern Nothotrichocera antarctica ( Edwards, 1923) and Jurassic Mailotrichocera jurassica Kalugina, 1985 ( Krzemińska 2001: fig. 20; Krzemińska, Krzemiński et Dahl 2009: fig. 19B).