Laciniomyia dilata, Kung & Brown, 2005

Kung, Giar-Ann & Brown, Brian V., 2005, Three unusually divergent new genera of phorid flies (Diptera: Phoridae) from Costa Rica, Zootaxa 1019 (1), pp. 43-52 : 44-45

Laciniomyia dilata new species

Figs 1–2 .

Description. Female (male unknown). Body length 1.42 to 1.48 mm. Head dorsoventrally flattened. Ocelli present. Frons brown, with anteromedial apex broadly rounded, not pointed. Frontal ratio 0.88. Median furrow of frons absent. All twelve frontal setae present (one pair each of dorsal and ventral interfrontal setae, dorsal and ventral fronto­orbital setae, inner vertical setae and post­ocellar setae). Dorsal fronto­orbital seta high, located near dorsal margin of eye, just below vertical seta. Ventral fronto­orbital seta near to eye margin. Two pairs of long, ventrally­directed supra­antennal setae present; setae approxi­ mately 1.5 times length and thickness of ventral interfrontal setae. Eyes oval, reduced, approximately 1.5 times size of flagellomere 1. Flagellomere 1 brown, elongate oval, with long yellow setae. Setae of apical third of flagellomere 1 at least twice as long as that in basal two­thirds. Arista apical, setose. Palpus dark brown, broad, long, approximately 1.3 times length of frons; dorsoventrally flattened, with concave dorsal face, color of concavity lighter than rest of palpus. Apical one­quarter of palpus with brush of approximately 30 to 40 long setae ( Fig. 2 ), length of setae approximately one­half length of palpus. Mouthparts whitish. Clypeus broad. Labrum elongate, thin. Labella elongate, thin, pointed. Oral cavity large, formed by greatly enlarged, protruding postoral expansion. Scutum brown, yellow anteromedially. Scutellum brown. Posterior pair of scutellar setae approximately 1.5 times length and thickness of anterior pair. Notopleural cleft absent. Anepisternum divided, and with longitudinal ridge on anterior portion. Anepisternum without setae. Wing length 1.30 to 1.38 mm. Mean costal length 0.31 wing length. Costal sector ratio 1.0:1. Costal setae long, apicalmost seta nearly as long as costa. R 2+3 absent. Halter yellow, elongate. Legs mostly yellow, tarsomeres brown. Forefemur with long, slightly thickened, widely spaced, posterodorsal to dorsal setae. Foretibia short, only slightly longer than onehalf length of forefemur. Foretarsomeres with long setae, tarsomere 1 with dense, long, ventral setae. Midtibia without differentiated pair of setae; with slightly thickened, longer, anterodorsal to dorsal setae along length. Hind femur apically dilated, expanded; base of femur narrow. Hind tibia with one longitudinal setal palisade. Hind tarsus tapered; tarsomere 1 extremely deep, laterally flattened, twice as deep as tarsomere 2. Dorsum of abdomen with long, thin, sparse setae. Abdominal tergites reduced, brown. Tergite 1 extremely short, narrowed. Tergite 2 same width as tergite 1, slightly tapering apically, not short. Tergite 3 approximately one­third width of tergite 2. Tergite 4 approximately two­thirds width tergite 3. Abdominal segment 4 with posterior gland opening, diameter of which only slightly smaller than width of tergite 4. Abdominal segments 5­6 lacking tergites. Abdominal segments 1–5 whitish to yellow, with long, thin, sparse setae; segment 6 brown with extremely long, sparse setae, setae as long as segment and extending beyond segment. Ovipositor mostly membranous, not parasitic­type.

Geographic distribution. Costa Rica.

Derivation of names. The generic name is from Latin lacina for fringe, referring to the apical brush of setae on the palpus. The species name is Latin for dilated, referring to the shape of the hind femur.

Holotype: ♀. COSTA RICA: Guanacaste: Estación Biologica Cacao , 10.93°N, 85.47°W, 10­11.vii.2000, J. Ashe, R. Brooks, Z. Falin, FIT, 1050 m [ LACM ENT 068947 ] ( LACM).

PARATYPE: 1♀, same data as holotype ( LACM) .

Figures

FIGURES 1–3. Laciniomyia dilata new species. 1. Abdomen, dorsal. 2. Body, lateral. 3. Laishania angustithorax new species. Body, lateral.