Subepona leahae, 2023

Groves, Lindsey & Squires, Richard L., 2023, Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover, PaleoBios 40 (10), pp. 1-52 : 12

SUBEPONA LEAHAE SQUIRES AND GROVES N. SP.

FIGS. 4O–Q

Cypraea species indeterminate. Squires, 1988. p. 12, fig. 23. Groves and Squires, 2021. p. 227.

Eocypraea (Eocypraea) castacensis ( Stewart, 1926 [1927]). Squires, 2000. p. 896.

Zoobank ID — LSID: urn:lsid:zoobank.org:act:363755AA-DB51-40D7-B996-A9EBA6EBBD85

Holotype and Type Locality— LACMIP 40374.79 , LACMIP Type 14936 (Figs. 4O–Q), length 29.5 mm, width 13.9 mm, height 15.1 mm. LACMIP Locality 7242 [= LACMIP 40374 ; ex CSUN Locality 374], Llajas Forma- tion, “Stewart bed,” Las Llajas Canyon , north side of Simi Valley , Ventura County, southern California.

Paratype — LACMIP 16178.1, LACMIP Type 7704, length 23.2 mm, width 17.1 mm, height 13.8 mm, LACMIP Locality 16178 [= LACMIP Locality 40984; ex CSUN Lo- cality 984) Lockwood Valley, Ventura County, California. This specimen was formerly hypotype LACMIP 7704 (of Squires, 1988: p. 12, fig. 23), of Eocypraea castacensis ( Stewart, 1926 [1927]).

Occurrence —“ Domengine Stage”: Juncal Formation?, Lockwood Valley , Ventura County, southern California ; Llajas Formation, Las Llajas Canyon (“Stewart bed,” ex CSUN Locality 374), Ventura County, California .

Etymology— Named for Leah Rose Regan, grand- daughter of R.L. Squires (who collected the type mate- rial).

Description— Shell medium size, moderately in- flated, aperture fairly straight except for moderate bend posteriorly, fossula smooth, teeth numerous and small on entire lengths of both lips, with those on the outer lip becoming slightly coarser near anterior end of shell. Posterior end of shell with outer lip projecting out over the inner lip. Anterior end with an elongate, linear pro- jection on each lip.

Remarks— The two known specimens of Subepona leahae have good preservation. The holotype is from the “Stewart bed” of the Llajas Formation, and the paratype is from the Juncal Formation?, Lockwood Valley, Ventura County, southern California.

Squires (1988: p. 12, fig. 23) reported a specimen (hypotype LACMIP 7704) of “ Cypraea species inde- terminate” [now identified as Subepona leahae ] from LACMIP Locality 40984 [ex CSUN Locality 984] in the Juncal Formation? in Lockwood Valley, Ventura County, southern California. This locality is in the upper 100 m of the Juncal Formation, based on work by Squires (2000: p. 896), who identified this same specimen as Eocypraea (Eocypraea) castacensis and revised its age to the younger “Domengine Stage,” instead of the previously reported ( Squires 1988) “Capay Stage.”

Subepona leahae differs from S.goedertorum by having much finer denticulation.