LUPONOVULA MANIOBRAENSIS

Groves, Lindsey & Squires, Richard L., 2023, Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover, PaleoBios 40 (10), pp. 1-52 : 33

LUPONOVULA MANIOBRAENSIS

(SQUIRES AND ADVOCATE, 1986) N. COMB.

FIGS. 9M–P

Eocypraea? maniobraensis Squires and Advocate, 1986 . pp. 856, 857, figs. 2.5, 2.6. Squires, 1991. p. 219, 220, 225; pl. 1, fig. 23.

Eocypraea (Eocypraea) maniobraensis Squires and Advocate. Groves, 1992 . p. 106 [not 1987 as in table 1]. Groves, 1993. p. 11 [not 1983 as in text]. Groves, 1997. p. 8 [as E. (E.) maniobraensis ]. Groves, 2011. p. 46 (table 1).

Eocypraea maniobraensis Squires and Advocate. Fehse, 2001 . p. 20. Groves and Squires, 2021. p. 231 (as Eocypraea ?). Fehse, 2021. p. 42.

Holotype and Type Locality— LACMIP10566.4 , LAC- MIP Type 10566 (ex UCLA 48431 ) (Figs. 9M–O), length 39.6 mm, width 22.8 mm, height 21.1. LACMIP Locality 23779 [ex UCLA Locality 3779; ex CSUN Locality 662], Maniobra Formation , Orocopia Mountains , Riverside County, southern California.

Referred Specimen—Hypotype (herein) LACMIP 40491.1 , LACMIP Type 14945 (Fig. 9P) , LACMIP Locality 40491 [ex CSUN Locality 491], Llajas Formation, is from a faulted area low in the stratigraphic section of this for- mation (see geologic map in Squires 1984: fig. 16) in Las Llajas Canyon , north side of Simi Valley , Ventura County, southern California [new information]. This Llajas For- mation specimen has been crushed but retains enough morphological features to identify it as L. maniobraensis .

Occurrence— Lower Eocene (“Capay Stage”), Llajas Formation, southwestern Santa Susana Mountains, Los Angeles County, southern California, Ventura County (new information) to Maniobra Formation, Orocopia Mountains, Riverside County, southern California ( Squires and Advocate 1986; Squires 1991).

Etymology— Originally named for the Eocene Manio- bra Formation, Riverside County, southern California.

Description— Shell size medium; very inflated; ovatepyriform. Shell widest medially. Base inflated. Spire in- volute. Aperture narrow and near right margin of shell; aperture elongate anteriorly and curved posteriorly. Aperture narrow and prominently curved. Outer lip nar- row, with numerous teeth that become stronger toward anterior of aperture. Inner lip with numerous small teeth.

Remarks— This is an uncommon species. The an- teriormost part of the holotype is missing. The dorsal view of this species is shown for the first time in Figure 4M. Luponovula maniobraensis most closely resembles L. oligovata ( Sacco 1894: pl. 3, fig. 25; Cossmann 1903: pl. 7, fig. 2) of late early Oligocene age (Latorfian) from the Liguria and Piedmont regions of Italy and L. normalis ( de Gregorio 1880: pl. 6, fig. 14) of middle Eocene age (Lutetian) from San Giovanni Illarione, Veneto, Italy. In comparison, Luponovula maniobraensis differs by having fewer teeth on its columellar lip, as well as having fewer and shorter teeth on the interior of its outer lip.