Liogluta Thomson, 1858

Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit & Ernst, Crystal, 2016, A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae), ZooKeys 573, pp. 217-256 : 219

Liogluta Thomson, 1858

Liogluta Thomson, 1858: 35. Type species Homalota umbonata Erichson, 1839, by monotypy. As valid genus: Lohse (1974); Lohse et al. (1990).

Anepsiota Casey, 1894: 321; Casey (1906: 335); as Atheta (Anepsiota) : Casey (1910: 12), Fenyes (1920: 203). Synonymized by Bernhauer and Scheerpeltz (1926: 656); Moore and Legner (1975: 350).

Athetota Casey, 1906: 334. Synonymized with Atheta (Anepsiota) by Fenyes (1920: 203); as synonym of Atheta (Liogluta) : Bernhauer and Scheerpeltz (1926: 656), Moore and Legner (1975: 350).

Hypnota Mulsant & Rey, 1873: 591. Synonymized with Atheta (Liogluta) by Fenyes (1920: 203); Bernhauer and Scheerpeltz (1926: 656); Moore and Legner (1975: 350).

Diagnosis.

Body length ranging from 2.8-5.4 mm, body narrowly subparallel (Figs 1, 8, 16, 23, 30, 35, 44, 50, 58, 65, 72, 79, 86, 90), moderately flattened; elytra and abdomen wider than head and pronotum; uniformly dark brown or reddish-brown with head and posterior abdomen dark brown to almost black; integument of forebody with strong meshed microsculpture; surface of elytra often granulose (Figs 50, 51); head with incomplete postocular carinae, postocular region of head long and in most species longer than diameter of eye; glossae split into two lobes; antennae long with articles VI-X subquadrate, slightly transverse or rarely slightly elongate; pronotum with hypomera fully visible in lateral view; pubescence of pronotum directed posteriad on midline of disc and posterolaterad to laterad on sides; mesocoxae narrowly separated, metasternal process short and acute; legs long, three basal articles of metatarsi highly elongate in most species; tarsal formula 5-5-5. Male. Apical margin of male tergite VIII often with broad, variably-shaped rectangular projection, with edge entire or bearing crenulation or variably shaped structures, with two lateral teeth and sometimes with additional median tooth; integument of disc often with broadly distributed asperate punctation near apex (Figs 3, 11, 18, 25, 38, 46, 53, 60, 67, 74, 81, 88, 92); median lobe of aedeagus simple with apical part variably shaped in lateral view (Figs 2, 9, 17, 24, 36, 45, 52, 59, 66, 73, 80, 87, 91). Female. Spermatheca with capsule narrowly club-shaped or tubular, with apical invagination moderate to deep, stem long, sinuate, looped or twisted posteriorly (Figs 7, 15, 22, 29, 34, 42, 43, 57, 64, 71, 78, 85, 96); sternite VIII with apical margin rounded or medially emarginate, antecostal suture arcuate, or slightly to distinctly sinuate (Figs 6, 14, 21, 28, 33, 41, 49, 56, 63, 70, 77, 84, 89, 95).

Distribution.

The genus is holarctic in distribution ( Smetana 2004). Seevers (1978) mentioned a few species from Africa and Jamaica but these records need verification.

Key to Canadian and Alaskan species of Liogluta