Mallinella annulipes ( Thorell, 1892 ) Dankittipakul & Jocqué & Singtripop, 2012

Dankittipakul, Pakawin, Jocqué, Rudy & Singtripop, Tippawan, 2012, Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from Southeast Asia (Araneae, Zodariidae) 3369, Zootaxa 3369 (1), pp. 1-327 : 164-165

Mallinella annulipes ( Thorell, 1892) nom. nov.

( Figs 712–713, 719–722, 724–725, 730, 733–734, 737–747)

Storena annulipes Thorell, 1892: 209 , description of juvenile, preoccupied in Storena . Workman 1896: 76, pl. 76, description of ♀

Storena cinctipes Simon, 1893: 430 , replacement name

Storena pseliphora Thorell, 1894: 5 , superfluous replacement name

Mallinella cinctipes, Song, Zhang & Li, 2002: 374 , transfer from Storena

New material. MALAYSIA, Pahang State: 1♂, 1 juvenile, Taman Negara, trail from Nusa Camp to Abai Waterfall (04°23'48.7"N, 102°25'50.0"E), 110 m, rainforest near stream, 16–17 June 2004, leg. P.J. Schwendinger ( MHNG, MAL –04/14) . Johor State: 1♀, Layang–Layang , July 1979 ( FJMC, coll. 7692) . SINGAPORE: 3♂, 1♀, Timah Hill , February 1988, leg. F. & J. Murphy ( MC, Coll. 15471) ; 1♂, Upper Peirce Reservoir , February 1988, leg. F. & J. Murphy ( FJMC, coll. 15443) ; 1♂, Upper Peirce Reservoir , March 1988, leg. F. & J. Murphy ( FJMC, coll. 13418) . INDONESIA, Singkep Island : 1♀, Batu Ampar Waterfall, ca. 10 km northwest of Dabo (00°29'30.9''S, 104°28'31.4''E), 80 m (rain forest along stream); 17–18 June 2001, leg. P. Schwendinger ( MHNG, SIM –01/04) .

Taxonomic remarks. Storena annulipes Thorell was originally placed in Storena and preoccupied with S. annulipes (L. Koch, 1867) . Mallinella annulipes nom. nov. is here removed from its replacement name with S. cinctipes Simon 1893 .

Diagnosis. Mallinella annulipes nom. nov. can be recognized by the color of the legs ( Fig. 103) which are provided with broad, dark brown distal and proximal annulations on femora and tibiae [hence the replacement specific epithet cinctipes (= banded legs) was proposed by Simon], in combination with an elevated dorsal hump on femora I–III in males. The male of this species is described by the first time. Males can be distinguished by the apically bifid embolus ( Fig. 739) and the banded legs; females by the narrowed, V-shaped epigynal plate and by the digitiform lateral borders projecting inwards ( Fig. 733).

Description. Male (here designated, MHNG, MAL–04/14). Total length 5.60; prosoma 3.44 long, 2.25 wide. Eye sizes and interdistances: AME 0.20, ALE 0.18, PME 0.16, PLE 0.18, AME–AME 0.10, AME–ALE 0.16, PME–PME 0.16, PME–PLE 0.30; MOQ: 0.46 long, 0.40 anterior width, 0.38 posterior width. Leg measurements: I 14.62 (3.41, 4.29, 4.18, 2.72), II 13.74 (3.20, 3.64, 3.56, 3.22), III 12.46 (3.00, 3.42, 3.52, 2.44), IV 16.90 (3.48, 4.64, 5.80, 2.94).

Coloration ( Figs 712, 719–722.). Carapace dark reddish brown. Sternum brown. Chelicerae dark brown. Legs yellowish, except coxae pale yellow; femora and tibiae with dark brown distal and proximal annulations. Opisthosoma sepia. Dorsal pattern: first and second pairs represented by pale round spots obliquely arranged; third and fourth pairs by transverse pale spots distinctly smaller than anterior pairs; fifth pair situated far apart, larger than third and fourth pairs, interspersed with a small triangular patch. Legs modified: femora I–III dorsally with elevated hump and stridulating files.

Palp ( Figs 730, 739–742). RTA flange-like, broad at base, gradually tapering towards bluntly pointed apex. TA rostrated, without distinct ridge; apico-prolateral fold elongated, sharply pointed, directed mesad. Tegular spine broad, terminally blunt. Embolic base with parallel margins, anterior membranous area almost indistinct. Embolus broad, with deep longitudinal groove running from proximal part to apex; mesal flange subterminally divided into two sharply pointed rami.

Female ( Singapore, MHNG). Total length 5.26; prosoma 3.23 long, 2.10 wide. Eye sizes and interdistances: AME 0.18, ALE 0.16, PME 0.16, PLE 0.16, AME–AME 0.08, AME–ALE 0.16, PME–PME 0.16, PME–PLE 0.26; MOQ: 0.42 long, 0.38 anterior width, 0.3368 posterior width. Leg measurements: I 13.72 (3.20, 4.02, 3.92, 2.56), II 12.71 (3.00, 3.42, 3.35, 2.39), III 11.66 (2.86, 3.22, 3.32, 2.28), IV 11.58 (3.26, 4.36, 5.44, 2.76).

Coloration ( Fig. 713). Carapace dark brown. Sternum brown. Chelicerae dark brown. Legs yellowish, except coxae pale yellow; femora and tibiae with dark brown distal annulations. Opisthosoma dark sepia. Dorsal pattern: first and second pairs represented by pale round spots obliquely arranged; third and fourth pairs by transverse pale spots distinctly smaller than anterior pairs; fifth pair situated far apart, larger than third and fourth pairs, interspersed with a small triangular patch.

Genitalia ( Figs 733–734, 737–738, 744–747). Epigynal plate narrowed, V-shaped. Lateral borders digitiform, terminally blunt. Insemination ducts slightly less than spermathecal length. Spermathecae short and truncated, provided with small apical hump in anterior view.

Natural history. Mallinella annulipes nom. nov. inhabits the forest floor of tropical lowland rainforests.

Distribution. West Malaysia, Singapore and Indonesia (Singkep Island).