Mallinella kritscheri, Dankittipakul & Jocqué & Singtripop, 2012

Dankittipakul, Pakawin, Jocqué, Rudy & Singtripop, Tippawan, 2012, Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from Southeast Asia (Araneae, Zodariidae) 3369, Zootaxa 3369 (1), pp. 1-327 : 129-130

Mallinella kritscheri sp. nov.

( Figs 537 , 553–554 , 567, 569 , 576–577 , 584–586 )

Storena sobria, Kritscher, 1957: 269 , fig. 28, description of ♀ (misidentification)

Material examined. Holotype: ♀, INDONESIA, Sumatra, West Sumatra Province: (at the foot of the Mt. Sigalang) Anai Valley (= Lambah Anai ), 6 km south of Padang Panjang (00º 28'38"S, 100º 21'14"E), 500 m, primary forest, 1 June 2006, leg. P.J. Schwendinger ( MHNG, Sum –06/03) . Paratypes: INDONESIA, Sumatra, North

Sumatra Province: 1♀, Gunung Leuser NP, north part around Bohorok rehabilitation center, sink C, primary rainforest on hills, 200 m, 28 1983, leaf litter, leg. Suh. Djojosudharmo ( RMHN); 1♀, 7 March 1983, leaf litter, leg. Suh. Djojosudharmo ( RMHN). Sumatra, Si Rambe: 1♀, female lacking genital + 1 slide preparation with text ‘ Internal genitaliaa & epigyne – Allotype!’ prep. Kritscher ; 1♀, 1 immature specimen ( NHMV, examined). Original label: / Sumatra, Si Rambe , leg. Modigliani. - Sammlung Reimoser ; Reimoser don. (A.-D. 15.X.1938), det. Inv. Nr. 433/. One male located in the same vial with these paratypes was correctly considered a male of M. sobria comb. nov. by Kritscher and that is confirmed by us.

Remarks. Note on the status of the allotype designated by Kritscher. Muttkowski (1910: 10) first introduced the term allotype in his ‘Catalog of the Odonata of Northern America’. According to Muttkowski (1910), allotype ‘need not be described by the proctologist; it can be contained in the original as well as in any subsequent description by other authors’. Unfortunately, most scientists were misled as to its original definition and purpose. According to the International Code of Zoological Nomenclature (2000), types are only those specimens which were designated as types in the original description (in this case, those studied and mentioned by Thorell). Therefore, the female indicated by Kritscher as ‘allotype’ of S. sobria is not a type but non-type material or additional material. It is here designated as a paratype of M. kritscheri sp. nov.

Diagnosis. This new species can be easily recognized by the epigynal plate provided with a longitudinal median septum ( Fig. 553 ), and by the elongated, ovoid spermathecae ( Fig. 554 ).

Etymology. This new species is named in honor of Professor E. Kritscher who first described the female (in S. sobria ).

Description. Female (holotype). Total length 7.62; prosoma 4.02 long, 3.10 wide. Eye sizes and interdistances: AME 0.18, ALE 0.12, PME 0.12, PLE 0.12, AME–AME 0.06, AME–ALE 0.18, PME–PME 0.12, PME–PLE 0.28; MOQ: 0.50 long, 0.42 anterior width, 0.38 posterior width. Leg measurements: I 17.80 (4.56, 4.70, 5.36, 3.27), II 15.24 (3.90, 3.90, 4.14, 3.21), III 13.72 (3.46, 4.30, 3.42, 2.52), IV 17.94 (4.60, 4.14, 5.90, 2.38).

Coloration ( Fig. 537 ). Carapace orange-brown. Sternum reddish brown. Chelicerae brown. Legs yellowish, except femora distally darker than other segments. Opisthosoma sepia, with three chevrons and two triangular bands longitudinally arranged.

Genitalia ( Figs 553–554, 556–557 , 584–586 ). Epigynal plate anchor-liked, provided with anterior median septum. Proximal part of spermathecae cylindrical, moderately long, with spherical posterior lateral swelling; distal part of spermathecae conical, divergent, elongated and ovoid in anterior view, internally with 5–6 coils.

Male. Unknown.

Natural history. This new species occurs syntopically with M. sobria comb. nov. on the floor of primary rainforests.

Distribution. Northern Sumatra, Indonesia.

Figures

FIGURES 535–538. 535. Mallinella sobria comb. nov., new material from Mt. Sigalang, Sumatra, male. 536. M. sobria comb. nov., new material from Mt. Sigalang, Sumatra, female. 537. M. kritscheri sp. nov., holotype. 538. M. caperata sp. nov., paratype. 535–538. Habitus, dorsal.

FIGURES 545–556. 545–548. Mallinella cirrhifera sp. nov., paratype. 549–550. M. caperata sp. nov., paratype. 551–552. M. sobria comb. nov., new material from Mt. Sigalang. 553–554. M. kritscheri sp. nov., holotype. 555–556. M. clavigera sp. nov., paratype. 545, 547, 549, 551, 553, 555. Epigyne. 546, 548, 550, 552, 554, 556. Internal genitalia, anterior.

FIGURES 565–569. 565. Mallinella sobria comb. nov., syntype (MCSN). 566, 568. M. sobria comb. nov., male from Mt. Sigalang (NHMV). 567, 569. M. kritscheri sp. nov., paratype (NHMV). 565–567. Habitus, dorsal. 568. Eye region, dorsal. 569. Epigyne.

FIGURES 570–577. 570–572. Mallinella sobria comb. nov., male from Mt. Sigalang (NHMV). 573–575. M. sobria comb. nov., holotype (MCSN). 576–577. M. kritscheri sp. nov., paratype (NHMV). 570. Male palp, ventral. 571, 575. Ditto, retrolateral. 572, 574. Ditto, prolateral. 573. Ditto, dorsoprolateral. 576. Internal genitalia, ventral. 577. Ditto, dorsal.

FIGURES 584–586. 584–586. Mallinella kritscheri sp. nov., holotype (RMHN). 584. Right spermatheca, dorsal. 585. Ditto, anterior. 586. Left spermatheca, lateral.