Deferunda Distant, 1912

Chen, Xiang-Sheng & He, Ting-Ting, 2010, Two new species of genus Deferunda Distant (Hemiptera: Fulgoromorpha: Achilidae) from southwest China, Zootaxa 2335, pp. 59-68 : 60

Deferunda Distant, 1912

( Figs 1–22 )

Majella Kirkaldy, 1906:421 preoccupied by Majella Ortmann 1893

Deferunda Distant, 1912:186 ; Fennah, 1950: 104; Chen et al., 1989: 57; Wang et al., 2008: 775. Okatropis Matsumura, 1914:272 , synonymised by Fennah 1950: 104.

Majellana Metcalf, 1948:63 , nom. nov. for Majella Kirkaldy , synonymised by Fennah 1950: 104.

Type species. Deferunda stigmatica Distant, 1912: 186 , by original designation.

Description. The distinctive characters used by Fennah (1950) and Chen, Yang and Wilson (1989) are modified as follows.

Head and thorax. Head with eyes ( Figs 1 , 13 ) distinctly narrower than pronotum (0.7:1). Vertex not declivous, produced before eyes two-thirds to a half length of vertex, disk strongly depressed, wider between basal angles than long in middle line (0.88–1.3:1), wider between basal angles than at apex (5:1), posterior margin subtruncate or slightly concave, anterior half of median carina obsolete, basal half prominent, lateral margins distinctly foliate, highly elevated, diverging posteriorly. Frons ( Figs 2 , 14 ) slightly convex in profile, longer in middle line than widest part (1.16–1.36:1), basal margin roundly convex or truncate one-fifth as wide as broadest part; median carina simple, with basal half obsolete, lateral margins strongly foliate basally, extending laterad beneath antennae, hence incurved into suture, disk of frons depressed at basal third, or apparently so on account of deeply foliate margins. Postclypeus shorter than frons in middle line about 0.50– 0.83:1, nearly straight in profile. Eyes concave beneath. Ocelli detached from eyes. Antennae subovate. Rostrum relatively short, reaching median trochanters, with apical and subapical segments equal in length. Pronotum shorter than vertex (0.52–0.64:1), almost as long behind eyes in middle line, disk slightly depressed between median and lateral carina, anterior margin angularly or rounded convex, lateral carinae diverging posteriorly, reaching hind margin, 1.6–2.0 times length of median line, each lateral lobe with four obsolete ridges. Mesonotum longer than vertex and pronotum combined (1.88–2.11:1). Forewing ( Figs 3 , 15 ) costal margin slightly convex, longer than wide (2.4–3.1:1), Sc+R forked in basal third of forewing, Cu1 forked level with union of claval veins, M forked level with node, Cu1 deeply convex distad of claval apex, almost reaching M, hence slightly detached; with a callus in costal cell, six apical areoles distad of stigma, apical part behind apex of clavus dropping and covering apex of abdomen. Spinal formula of hind leg 8–7(8)–6(5).

Male genitalia. Anal segment in dorsal view ( Figs 5 , 16 ) rounded with apical margin truncate or emarginated, anal style moderately long. Pygofer in profile ( Figs 6 , 17 ) distinctly shorter dorsally than ventrally, anterior margin concave, posterior margin convex caudad, pygofer in ventral view ( Figs 7 , 18 ) with a pair of medioventral processes. Aedeagus ( Figs 11, 12 , 21, 22 ) with phallobase dividing into four lobes at apex, ventral lobe usually cleft at apex medially. Phallic appendages slender, very long, tubular, tapering apically. Genital styles ( Figs 8-10 , 19, 20 ) narrow at base, and widening apically, each nearly triangular, outer margin usually with one or two teeth apically and subapically. Connective ( Figs 8 , 19 ) very long.

Host plant. Unknown.

Distribution. Oriental, Palaearctic and Australian regions ( Fig. 23 ).

Figures

FIGURE 23. Geographic distribution of Deferunda species: D. acuminata (); D. albomaculata (); D. diana (); D. incompta (); D. lineola (); D. majella (); D. philippina (); D. qiana (); D. rubrostigmata (); D. stigmatica (); D. striata (); D. trimaculata (); D. truncata ().

FIGURES 1 – 12. Deferunda diana (1) Head and thorax, dorsal view; (2) Frons and clypeus; (32) Forewing; (4) Hindwing; (5) Anal segment, dorsal view; (6) Pygofer, lateral view; (7) Pygofer, ventral view; (8) Genital styles and connective, ventral view; (9) Genital style, lateral view; (10) Genital style, ventral view; (11) Aedeagus, ventral view; (12) Aedeagus, dorsal view. Scale bars = 0.5 mm (Figs 1, 2), 1.0 mm (Figs 3 – 4), 0.2 mm (Figs 5 – 12)

FIGURES 13 – 22. Deferunda qiana (13) Head and thorax, dorsal view; (14) Frons and clypeus; (15) Forewing; (16) Anal segment, dorsal view; (17) Pygofer, lateral view; (18) Pygofer, ventral view; (19) Genital styles and connective, ventral view; (20) Genital style, lateral view; (21) Aedeagus, ventral view; (22) Aedeagus, lateral view. Scale bars = 0.5 mm (Figs 13, 14), 1.0 mm (Fig. 15), 0.2 mm (Figs 16 – 22)