Atopocelyphus Gaimari

Gaimari, Stephen D., 2017, The dipteran family Celyphidae in the New World, with discussion of and key to world genera (Insecta, Diptera), ZooKeys 711, pp. 113-130 : 115

Atopocelyphus Gaimari gen. n.

Type species.

Celyphus ruficollis Macquart, 1844, by present designation.

Etymology.

From Greek, Atopos , meaning out of place, combined with the genus name Celyphus , referring to the unexpected occurrence of this taxon in the New World; masculine.

Diagnosis.

This genus differs from all other Celyphidae in having an elongate first flagellomere with a subbasal, plumose arista (Fig. 13), and in having abdominal tergites 5 and 6 each subdivided or creased medially with a strong triangular notch along each posterior edge in both sexes (Figs 18, 20).

Remarks.

The other celyphid genera have a much shorter first flagellomere with a subapical arista that is pubescent and often expanded and leaf-shaped in the basal 1/3 (see Fig. 29). The abdominal tergites are sometimes subdivided (i.e., in Spaniocelyphus ), but this is always tripartite, with a central section and two lateral sections (Fig. 31); otherwise, the tergites are undivided (Fig. 28). With regards to other dipteran families in the Neotropics with superficially similar genera, Celypholauxania Hendel ( Lauxaniidae ) and Peltopsilopa Hendel ( Ephydridae ) share a characteristically enlarged scutellum, although none to the extent of the Celyphidae . One of the species currently in Peltopsilopa had been originally described as a species of Celyphus ( Savaris et al. 2016), and other genera (outside the New World) had also been originally described as celyphids, such as Afrocelyphus Vanschuytbroek, now considered a junior synonym of Nomba Walker ( Chloropidae ).