Amphineurus (Rhamphoneurus) caleuchus, Santos & Santos & Ribeiro, 2022

Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha, 2022, Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae), Arthropod Systematics & amp; Phylogeny 80, pp. 439-494 : 439

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Amphineurus (Rhamphoneurus) caleuchus

sp. nov.

3.3.7. Amphineurus (Rhamphoneurus) caleuchus sp. nov.

Material examined.

Holotype: ♂, Chile, Chiloé Is. Ancud [41°52′S 73°48′W], 23-I-1932, Peña (USNM)*. Paratypes: 1 ♀, Chile, Chiloé Is., Aucar [42°9′S 73°30′W], 6/ 15-I-1952, Peña (USNM); 2 ♂, Chile, Chiloé Is., Chaiten [42°58′S 72°32′W], 5/ 8-II-1954, Peña (USNM)*. Additional material: CHILE. 1 [sex unknown], Chiloé Is., Aucar [42°9′S 73°30′W], 6/ 15-I-1952, Peña (USNM); 1 [sex unknown], Chiloé Is., Chaiten [42°58′S 72°32′W], 5-II-1954, Peña (USNM) GoogleMaps ; 1 ♂, Chiloé Is. Ancud [41°52′S 73°48′W], 18/ 19-XII-1926, F. & M. Edwards (USNM)*; 1 ♂, Chiloé Is., Aucar [42°9′S 73°30′W], 6/ 15-I-1952, Peña (USNM) GoogleMaps *; 1 ♂, Arauco, Nahuelbuta [37°46′S 72°59′W], 5/ 10-II-1953, Peña (USNM) [slide mounted specimen prepared by the authors] GoogleMaps .


The word Amphineurus caleuchus refers to the Chilean myth of Caleuche, a sentient ghost ship of the Chiloé islands ( Febles 2009). The species is named after this myth due to holotype and the majority of specimens being from the Chiloé islands.


The species is distinguished by a darkened distal quarter of the wing, R2+3+4 similar to R2+3, and M1+2 shorter than the basal deflection of M1. Furthermore, the species is characterized by the distinctive clasper of gonostylus with setae on medial branch, curved lateral branch and male tergite IX with a very shallow notch.


Wing length 6.50 mm, width 2.05 mm. - Coloration: General coloration dark brown. Rostrum and palpus dark brown. First segments of antenna yellow, remainding segments brownish-yellow. Head brown. Eyes black. Thorax yellowish-brown, pleura yellowish brown. Scutum with three dark brown stripes. Halter pale with knob ochreous. Coxae yellow and legs brown. Abdomen dark brown. - Head (dorsal view Fig. 14B View Figure 14 , ventral view Fig. 14A View Figure 14 ): Rostrum bifurcated with short appendices; first palpal segment shorter than terminal segment; scape as thick as pedicel; terminal flagellomere as long as penultimate; antenna with long setae. - Thorax (lateral view Fig. 14E View Figure 14 , dorsal view Fig. 14F View Figure 14 ): Thorax with short setae, some organized in rows on prescutum. Division line of laterotergite almost vertical; anatergite shorter than katatergite. Halter with dilated knob. Wing (Fig. 14D View Figure 14 ) almost clear of markings, except for faint markings on R2 and tip of R1; distal quarter of wing darkened; veins near fork of bM (Fig. 14C View Figure 14 ) faded; M1+2 shorter than basal deflection of M1; R2+3+4 as long as R2+3. - Female terminalia (Fig. 14H View Figure 14 ): Female tergite IX longer and narrower than tergite X; cercus thin and bent at right angle. - Male terminalia (Fig. 14G View Figure 14 ): Male tergite IX with sinuose posterior margin. Ventral branch of gonocoxite slightly inclined laterally, with long setae on lateral margin. Dorsal branch of gonocoxite reduced. Lobe of gonostylus with rounded lobule; lobule longer than wide, shorter than half-length of lobe of gonostylus, and narrower than stem. Clasper of gonostylus with differently shaped branches: lateral branch straightened, slightly bent towards tip; medial branch curved, club-shaped with concentration of setae. Mesal lobes symmetrical, rounded at posterior margin with bump on side. Sheath of aedeagus angulated with two bends; darkened almost along entire length.


Specimens of this new species were previously identified by C.P. Alexander as A. (R.) nothofagetorum Alexander. This species resembles A. (R.) stigmaticus sp. nov. but differs mainly in markings on the sheath of aedeagus, length of R2+3+4, and shape of lobe of gonostylus, cercus and mediotergite.