Calamagrostis scabrescens Griseb., Nachr. Koenigl . Ges. Wiss. Georg-Augusts-Univ. 3: 79. (1868)

Prasad, Dileshwar, Kumar, Ravindra, Jaiswal, Shubham, Yadav, Rekha, Tiwari, Smita & Agnihotri, Priyanka, 2022, An update on the taxonomy of Calamagrostis nagarum (Bor) G. Singh and its allies (Poaceae, Agrostidinae): morphometrics and micro-morphology, PhytoKeys 212, pp. 135-155 : 135

2. Calamagrostis scabrescens Griseb., Nachr. Koenigl. Ges. Wiss. Georg-Augusts-Univ. 3: 79. (1868)

C. scabrescens var. humilis Griseb. Nachr. Königl. Ges. Wiss. Georg-Augusts-Univ. 3: 79 (1868). Lectotype (designated here):-India. Sikkim, Lachen, 11,000 ft. [3352.8 m], [D. nr. 9], 3 July 1849, J.D. Hooker s.n. (K: K000838368, digital image!). (Image available at https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.k000032368).

Lectotype

(designated here). -India. Sikkim, Lachen, 12,000 ft [3600 m], [ Deyeuxia scabrescens Munr.], 3 August 1849, J.D. Hooker s.n. (K: K000838333, digital image!). (Image available at https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.k000838333)

Note.

Grisebach (1868) described Calamagrostis scabrescens with three varieties viz. (α) var. Calamagrostis scabrescens scabrescens Griseb., (β) var. Calamagrostis scabrescens elatior Griseb. and (γ) var. Calamagrostis scabrescens humilis Griseb. based on Hooker’s gatherings which were housed in Kew Herbarium. The name C. scabrescens (≡var. Calamagrostis scabrescens scabrescens ) was given for H. [ Hooker’s]: D. [ Deyeuxia ] Deyeuxia scabrescens Munr., characterized by ciliated glumes along the margin. The var. Deyeuxia scabrescens elatior was proposed for H. [ Hooker’s]: D. [ Deyeuxia ] nr. 7, in which glumes are not ciliated along the margin, panicle sub-violet and about 30 cm long. Var. Deyeuxia humilis was proposed for the H. [ Hooker’s]: D. [ Deyeuxia ] nr. 9, which differs from the above by having non ciliated glumes, panicle narrow and greenish, and ligule short, truncate or obtuse. Later, var. Deyeuxia humilis humilis was synonymized with C. scabrescens , while var. Calamagrostis scabrescens elatior raised to rank of species as C. elatior (Griseb.) A. Camus ( Camus 1928; Bor 1960).

While searching for Hooker’s specimen(s) belonging to the name C. scabrescens in various herbaria we traced six specimens, two at K (K000838333 and K000838334), one specimen at BM (BM000573477), CAL (0000004002), MPU (MPU027066), and W (W1889-0241775) each. All the specimens were collected from the different localities of Sikkim Himalaya and have the annotation "H.: Deyeuxia scabrescens Munr." The specimens at K, W, and CAL were verified as type materials by Beata Paszko. We have examined all the specimens including those which are housed in MPU and BM and determined as type material of C. scabrescens .

At CAL, the specimen, CAL0000002402, was stored under the type materials of C. scabrescens , without annotation of " Deyeuxia scabrescens Munr." on the sheet. This specimen was identified as the type of Deyeuxia filiformis senso. Hook. f., but later identified as C. scabrescens by Sunanda Bhattacharya, Botanical Survey of India. According to Paszko (2012a), it belongs to Deyeuxia filiformis senso. Hook. f. of C. lahulensis - C. scabrescens complex. Therefore, we excluded it from the type materials.

Since Grisebach examined Hooker’s specimens housed at K for his new species C. scabrescens , the specimen with barcode K000838333 (left-hand side) is designated here as lectotype for the name C. scabrescens as per Art. 9.3 of the ICN ( Turland et al. 2018), because of its good preservation with complete plants including inflorescence and spikelet and illustration of ciliated glumes on the sheet.

In addition to this, for the var. Calamagrostis scabrescens humilis we have traced another two specimens at K (K000838352 and K000032368) and one specimen at W (W0026817) which belong to the type materials of var. Calamagrostis scabrescens humilis . All of them bear the annotation "H.: D. nr. 9", and thus should be considered as syntypes (Art. 9.6 of the ICN; Turland et al. 2018). The specimen with barcode K000838368 is designated here as the lectotype for the name C. scabrescens var. humilis following Art. 9.3 of the ICN ( Turland et al. 2018), as the specimen is well-preserved and also morphologically complete with roots, inflorescences, and spikelets.