Bembidion (Zemetallina) wanakense Lindroth, 1976

Larochelle, André, Zeperyphodes, Broun & Larivière, Marie-Claude, 2015, Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini), Insecta Mundi 2015 (415), pp. 1-78 : 46-47

Bembidion (Zemetallina) wanakense Lindroth, 1976

Fig. 36 , 69 , 106

Bembidion (Zemetallina) wanakense Lindroth, 1976: 189 . Type locality: Matukituki River, West Branch, Northwest of Wanaka, OL.

Description. Body length 3.7–4.5 mm. Black; antennal segments 1–3 usually pale rufous (rarely dark rufous; infuscated above), segments 4–11 dark rufous; legs usually dark rufous (rarely piceous or black), tarsi infuscated. Microsculpture very strong, isodiametric. Shiny, with strong metallic lustre (aeneous). Forebody moderately wide in comparison to elytra. Head. Antennae filiform, moderately long (reaching about elytral shoulders). Thorax. Pronotum moderately convex and wide; sides strongly rounded anteriorly, moderately sinuate posteriorly; posterolateral angles rectangular, somewhat obtuse at tip; laterobasal foveae moderately deep, linear, moderately long, not reaching basal margin. Epipleura (in dorsal view) not exposed in front of posterolateral angles. Elytra. Moderately convex, subelongate, moderately wide, widest about middle. Shoulders moderately rounded. Sides subparallel. Scutellar striole consisting of a row of punctures. Striae 2–6 incomplete; striae 1–4 moderately deep, 5–8 shallow; striae moderately coarsely punctate; stria 7 obsolete, incomplete, with or without rudimentary punctures. Intervals 1–4 slightly convex, 5–8 depressed. Apical striole present, deep, connected to stria 5; preapical setiferous puncture not isolated. Abdomen. Last visible sternum (sternum VII) of female with four long ambulatory setae only. Aedeagus. Lateral view ( Fig. 69 ): strongly arcuate, wide; base slightly concave dorsally; middle strongly convex dorsally, moderately concave ventrally; apex subtriangular, straight dorsally, slightly concave ventrally, with extreme tip wide and short.

Material examined. 244 specimens ( CMNH, JNNZ, LUNZ, MONZ, NZAC).

Geographic distribution ( Fig. 106 ). North Island: WA. South Island: BR, CO, DN, FD, KA, MC, MK, NC, NN, OL, SC, WD.

Ecology. Lowland, montane. Fossorial. River banks and beds, at a certain distance (1–20 m) from water. Open ground; moist or wet, sandy, bare or sparsely vegetated (e.g., Juncus ) soil, sometimes mixed with silt. Nocturnal; hides during the day usually in the sand, sometimes at the base of plants and under fallen leaves. Gregarious.

Biology. Seasonality: September–May. Tenerals: February–March. Occasionally infested with fungi (Laboulbeniales). Defence mechanism: when alarmed, the adult escapes by running.

Dispersal power. Macropterous, probably capable of flight. Fast runner. Vagility likely favoured by flight capacity.

Collecting techniques. Pouring water over the ground; treading the soil with the feet; inspecting the base of plants; raking fallen leaves.

Reference. Larochelle and Larivière 2001: 83 (catalogue; biology, dispersal power, ecology, geographic distribution, reference).

Remark. Populations of Bembidion wanakense from the southern South Island (CO, DN, MK) sometimes have darker antennae and legs (antennal segment 1 piceous and tibiae black or piceous).

Figures

Figures 33–36. Dorsal habitus. 33) Bembidion bullerense new species. 34) B. hokitikense. 35) B. chalceipes. 36) B. wanakense. Scale line = 1 mm.

Figures 61–69. Aedeagi, lateral. 61) Bembidion parviceps. 62) B. waiho new species. 63) B. urewerense. 64) B. mangamuka new species. 65) B. solitarium. 66) B. bullerense new species. 67) B. hokitikense. 68) B. chalceipes. 69) B. wanakense.

Figures 101–106. Species distribution maps. 101) Bembidion tillyardi. 102) B. townsendi. 103) B. urewerense. 104) B. waiho new species. 105) B. waimarama new species. 106) B. wanakense.