Pentodontini

López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492 : 456

Key to the genera of adult Pentodontini from Colombia

1 Apical maxillary palpomere triangular, apically expanded ( Fig. 43 ).............................. Hylobothynus Ohaus

- Apical maxillary palpomere cylindrical.................................................................... 2

2 Clypeus triangular, acute, directed upward. Pronotum with small, apical tubercle and apical fovea large, deep, U-shaped............................................................................................ Oxyligyrus Arrow

- Clypeus truncate or narrowed towards apex, not strongly directed upward. Pronotum with tubercle and fovea variable, but fovea never U-shaped.................................................................................. 3

3 Clypeus trapezoidal, apex lacking teeth.................................................................... 4

- Clypeal shape variable, apex bidentate..................................................................... 6

4 Frontoclypeal suture present and strongly impressed, sometimes interrupted at middle; head without frontal tubercles. Size small (11–16 mm)..................................................................................... 5

- Frontoclypeal suture absent; head with a frontal tubercle near each eye, as an elongate horn in males and shorter in females. Size larger (20–30 mm)...................................................................... Pucaya Ohaus

5 Pronotum with 2 apical tubercles (easier to see in lateral view)..................................... Parapucaya Prell

- Pronotum without apical tubercles........................................... Denhezia Dechambre (incertae sedis)

6 Mandibles without teeth or only slightly sinuate (with acute apex and a basal lobe). Size small (13–15 mm).. Euetheola Bates

- Mandibles bidentate or tridentate. Size larger (usually 19–30 mm, rarely <19 mm).................................. 7

7 Mandibles tridentate, with sharp, triangular teeth. Clypeus strongly narrowed toward apex. Most species with apical pronotal fovea deep and wide (transverse diameter wider than head width).................................... Bothynus Hope

- Mandibles bidentate, with 2 apical teeth and a basal lobe, if tridentate then pronotum without or with inconspicuous fovea or tubercle. Clypeus narrowed towards apex or trapezoidal. Subapical pronotal fovea narrower (transverse diameter not wider than head width), or absent................................................................ Tomarus Erichson

Figures

FIGURES 40 – 48. Hylobothynus columbianus (holotype): 40) habitus in dorsal view, 41) parameres in frontal view, 42) parameres in lateral view, 43) maxillary apical palpomere, 44) habitus in lateral view. Hylobothynus obesus (holotype): 45) habitus in dorsal view, 46) parameres in frontal view, 47) parameres in lateral view, 48) locality records of Hylobothynus in Colombia.