Askola Peters

Domínguez, Eduardo, Molineri, Carlos & Mariano, Rodolfo, 2009, Revision of the South American species of Hagenulopsis Ulmer and Askola Peters (Ephemeroptera: Leptophlebiidae) with description of six new species, Zootaxa 2142 (1), pp. 29-44 : 31

Askola Peters

Askola Peters, 1969: 253 . (Type-species: Askola froehlichi Peters ).

Discussion: Askola can be distinguished from the other genera of Leptophlebiidae by the following combination of characters, in the imago: 1) hind wings absent; 2) MA fork slightly asymmetrical, MP fork symmetrical; 3) vein ICu1 of fore wings joined at base to vein CuA; 4) claws of a pair dissimilar, one apically hooked and the other blunt; 5) penes divided, tubular, slender, apically pointed, and without appendages or spines ( Figs. 28–36 ); and 6) female with egg guide prolonged to middle of abdominal sternum IX and 7) female sternum IX deeply cleft apically. In the nymph: 1) hind wing pads absent; 2) anteromedian emargination of labrum with two small denticles; 3) denticles of tarsal claws progressively larger apically; 4) dorsal and ventral portions of abdominal gills I–VII equal and plate-like, with the apical half of each plate bordered with finger-like processes; 5) posterolateral spines present on abdominal segments V–IX or VI–IX.

The generic description of the male imago needs to be emended with the following characters to include the variations reported for the new species: 1) eyes separated ( Fig 8 ) or meeting ( Figs. 2, 6 and 11 ) on meson of head; 2) wings hyaline ( Fig 27 ) or spotted ( Figs. 1 , 25, 26 ), costal cross veins basad to bullae absent ( Fig. 27 ) or present ( Figs. 25–26 ); 3) penes extending ( Figs. 28, 34 ) or not ( Fig 31 ) beyond posterior margin of styliger plate.

Figures

FIGURES 25–36. Askola. Figs 25–27, male fore wing: 25, Askola emmerichi sp.nov.; 26, Askola paprockii sp. nov.; 27, Askola cipoensis sp. nov. Figs 28–30, Askola emmerichi sp.nov.: 28, genitalia, ventral view; 29, genitalia, lateral view; 30, penes, detail. Figs 31–33, Askola paprockii sp. nov.: 31, genitalia, ventral view; 32, genitalia, lateral view; 33, penes, detail. Figs 34–36, Askola cipoensis sp. nov.: 34, genitalia, ventral view; 35, genitalia, lateral view; 36, penes, detail.

FIGURES 7–12. Askola. Figs 7–9, Askola cipoensis sp. nov., male imago. 7, general aspect, lateral view; 8, head, dorsal view; 9, head, lateral view. Figs 10–12, A. froehlichi, male imago. 10, general aspect, lateral view; 11, head, dorsal view; 12, head, lateral view.

FIGURES 1–6. Askola. Figs 1–3, Askola emmerichi, sp. nov., male imago. 1, general aspect, lateral view; 2, head, dorsal view; 3, head, lateral view. Figs 4–6, Askola paprockii sp. nov., male imago. 4, body, dorsal view; 5, idem, ventral view; 6, head, dorsal view.