Hyleoglomeris insularis Golovatch, 2013

Golovatch, Sergei, 2013, Three new species of the millipede genus Hyleoglomeris Verhoeff, 1910 from the Aegean region of Greece (Diplopoda, Glomerida, Glomeridae), Biodiversity Data Journal 1, pp. 1000-1000 : 1000

Classification: Glomeridae Rank: SpeciesType of treatment: New taxonextantHabitat: terrestrialRoot classification: 8

Hyleoglomeris insularis Golovatch, 2013 sp. n.

Materials

Type status: Holotype. Occurrence: recordedBy: P. Beron; sex: 1 male; Location: island: Kálimnos; country: Greece; verbatimLocality: village Scalia, Cave Scalia; Event: eventDate: 1987-05-04; Record Level: institutionCode: NMNHS Type status: Paratype. Occurrence: recordedBy: P. Beron; sex: 2 males, 2 females; Location: island: Kálimnos; country: Greece; verbatimLocality: village Scalia, Cave Scalia; Event: eventDate: 1987-05-04; Record Level: institutionCode: NMNHS Type status: Paratype. Occurrence: recordedBy: P. Beron; sex: 1 male, 1 female; Location: island: Kálimnos; country: Greece; verbatimLocality: village Scalia, Cave Scalia; Event: eventDate: 1987-05-04; Record Level: institutionCode: ZMUM

Description

Length of holotype ca 6.0 mm, width (maximum on tergum 2) ca 3.0 mm; length of paratypes ca 6.0-6.2 mm, width on tergum 2 ca 3.0-3.1 mm, or 6.2-7.5 and 3.2-3.4 mm in males and females, respectively. Body nearly entirely pallid (Fig. 5), only dorsal side of head retaining a faint to mediocre, rather uniform brownish coloration often growing a little darker on antennae and pale grey to nearly blackish around ocelli (Fig. 5b).

Ocelli 6+1 or perhaps 7+1, convex, completely translucid, but mostly clearly discernible due to an infuscated nearby background (Fig. 5b). Tömösváry’s organ pallid, transverse-oval, ca 1.4-1.5 times wider than long. Antennomere 6 long, ca 2.3-2.4 times as long as high.

Collum with two transverse striae. Tergum 2 with a rather broad hyposchism extending considerably behind caudal tergal margin (Fig. 5a); 4-5 superficial transverse striae, 1-2 starting below schism, remaining 2-3 above it, with three (never last one from below) crossing the dorsum. Male anal shield regularly rounded at caudal margin.

Male leg 17 (Fig. 6a) with a rather low, regularly rounded, outer coxal lobe; telopodite 3-segmented, tarsus with two strong apical spines.

Male leg 18 (Fig. 6b) with a narrow syncoxital notch; telopodite 4-segmented, tarsus with one apical spine.

Telopods (Fig. 6c) with a high, rounded, clearly emarginate, central syncoxital lobe flanked by two setose horns, each latter only slightly higher than central lobe and crowned by a minute, elongate, acute, membranous lobule devoid of adjacent structures. Only prefemur micropapillate laterally. Caudomedial femoral process prominent, directed distomedially at ca 100 º to femur, mostly strongly chitinized, only apically with a small membranous sac, but devoid of any chitinized lobe. Caudomedial process of tibia evident, sac-shaped, membranous, with an evident, rounded tubercle on caudal face at base. Tarsus rather modestly curved, subacuminate apically.

Diagnosis

Differs from congeners in a partly unpigmented body with only the head retaining some pigment, coupled with a long antennomere 6 which is ca 2.3-2.4 times as long as high, as well as by a rather broad hyposchism produced considerably behind the caudal margin of tergum 2, and only 4-5 transverse striae, of which three cross the dorsum on tergum 2.

Etymology

To emphasize the provenance from an island. An adjective.

Taxon discussion

Due to such a troglomorphic feature as a nearly completely unpigmented body, this species may well prove to be a troglobite. This cave on Kalimnos is known to support at least one more endemic troglobite, the woodlouse Cordioniscus kalimnosi Andreev, 1997 ( Isopoda , Oniscidea, Styloniscidae ) ( Schmalfuss 2003).