Ptomaphaminus layangensis Schilthuizen & Perreau

Schilthuizen, Menno, Perreau, Michel & Njunjic, Iva, 2018, A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae), ZooKeys 777, pp. 57-108 : 89-91

Ptomaphaminus layangensis Schilthuizen & Perreau sp. n. Figures 11c, 12s, t, 13o, p

Ptomaphaginus sp. n. bryanti complex Merckx et al., 2015: extended data fig. 6.

Material.

Holotype: Malaysia, Sabah, Kinabalu Park, Layang-Layang, 2750 m elev., 6°02.748'N, 116°33.632'E, 13-18.ix.2012 (leg. M. Schilthuizen, Crocker Range / Kinabalu Expedition, RMNH.INS.555619), 1 male. Paratypes: Sabah. Kinabalu Park, Layang-Layang, 6°02.748'N, 116°33.632'E, 2750 m elev., 13-18.ix.2012 (leg. M. Schilthuizen, Crocker Range / Kinabalu Expedition, CMPR, RMNH, incl. RMNH.INS.555618-555622), 11 paratypes.

Description.

Length: 1.6-2.5 mm. General colour dark brown; the tarsi and two first antennomeres lighter. Winged. Eyes well developed. Pronotum 1.6 times as wide as long. Elytra 1.45 times as long as wide. Sutural angle of female elytra rounded, without noticeable sexual dimorphism. Male protarsi 0.75 times as wide as the protibia. Spiculum gastrale of the male genital segment short, moderately dilated at the apex (Figure 13o). Aedeagus 5.4 times shorter than the body length, the sides regularly arcuate in dorsal view (Figure 12s), ventrally straight and dorsally obtusely angular in lateral view. Apex of the median lobe with a ventrally deflexed, arcuate hook in lateral view (Figure 12t). Six lateroventral preapical setae and one lateroapical seta on each side. Internal stylus of the endophallus significantly sinuate. Female genital segment with reduced gonocoxites. Spermiduct long and helical (Figure 13p). Apex of spermatheca rounded, without apical sclerotised plate.

Differential diagnosis.

Normal-sized species with unreduced eyes and non-spiniform female elytra. Aedeagus with a long, retroverted apical hook. Distinguishable from P. alabensis by the anteriorly widened spiculum gastrale and the broader apex of the median lobe of the aedeagus. Females are distinguishable from P. nanus by their larger size.

DNA barcodes.

COI barcodes are available in BOLD for the holotype, RMNH.INS.555619, as well as for the paratypes RMNH.INS.555618 and 555620-555622, jointly forming the BIN ACK0070.

Habitat and distribution.

Known from high altitude (above 2000 m) on Gunung Kinabalu, Sabah, Malaysia.

Etymology.

Named for the type locality on Gunung Kinabalu.