Goniurosaurus huuliensis Orlov, Ryabov, Nguyen, Nguyen & Ho, 2008

Ngo, Hai Ngoc, Nguyen, Huy Quoc, Tran, Hieu Minh, Ngo, Hanh Thi, Le, Minh Duc, Gewiss, Laurenz Rafael, Schingen-Khan, Mona van, Nguyen, Truong Quang & Ziegler, Thomas, 2021, A morphological and molecular review of the genus Goniurosaurus, including an identification key, European Journal of Taxonomy 751 (1), pp. 38-67 : 47-49

Goniurosaurus huuliensis Orlov, Ryabov, Nguyen, Nguyen & Ho, 2008

Fig. 3

Diagnosis

Body splayed and gracile, SVL 97.2–134.6 mm; external nares bordered by 6–8 nasal scales; supraorbital region with a row of enlarged tubercles; outer surface of upper eyelid composed of granular scales, about one half the size of those on top of head and without enlarged tubercles; internasal 1 or 2 (rarely absent); supralabials 9–12; infralabials 9–12; preorbital scales 14–20; eyelid fringe scales 51–59; postmentals 2–4; gular region below lower jaws with enlarged tubercles; paravertebral tubercles 31–37; scale rows around midbody 118–130, granular scales surrounding tubercles 11–13; axillary pockets deep; subdigital lamellae under fourth toe 21–25; precloacal pores in males 25–30; iris reddish brown; dorsal ground color of head, body and limbs dark brown, without small dark brown blotches (dark blotches present only on lower zone of flanks); nuchal loop thin, posteriorly protracted (in V-shape); dorsal body bands between limb insertions 3, thin, immaculate yellow; gular region with brown spots (modified after Orlov et al. 2008; Nguyen 2011).

Description (Supp. file 1: Table S2)

Body splayed and gracile, adult males SVL 97.2–132.2 mm (mean ± SE: 118.9± 1.4 mm, n =32), TaL 36.7–108.6 mm (72.7 ± 3.7 mm), adult females SVL 97.4–134.6 mm (121.1 ± 1.2 mm, n =46), TaL 48.4–106.7 mm (78.2 ± 2.7 mm), juveniles SVL 73.5–74.4 mm (n =2), TaL 62.5–62.7 mm (Supp. file 1: Table S2); head triangular, wider than neck, covered by uniform granular scales interspersed with tubercles in temporal and occipital regions; scales on rostrum slightly larger and flatter; enlarged supraorbital tubercles in a conspicuous row; middorsal portion of rostral partially sutured dorsomedially, bordered laterally by first supralabial on each side, dorsolaterally by prenasal on each side, and dorsally by internasal and two supranasals; internasal 1–3 (rarely absent); external nares bordered by 5–7 nasals (prenasal, anterior and posterior supranasals, 2 slightly enlarged postnasal, and 1 or 2 granular scales); preorbital scales 14–20; supralabials 9–12, grading into granular scales posteriorly; infralabials 9–12; eyes large, pupils vertical; eyelid fringe scales 51–59, those of upper eyelid slightly enlarged; outer surface of upper eyelid composed of granular scales of about one half the size of those on top of head, without enlarged tubercles; fold of skin originating from suborbital region extends posteroventrally across angle of jaw; external auditory meatus elliptical with long axis directed dorsoventrally; tympanum deeply recessed; mental triangular, bordered laterally by first infralabial on each side and posteriorly by 2–4 postmentals; postmentals bordered by 7–10 gular scales; gular region below lower jaws with enlarged tubercles; gular scales juxtaposed and granular, grading posteriorly into flat hexagonal pectoral scales and even larger ventral scales.

Neck narrower than body, covered with uniform granular scales interspersed with several sharply pointed conical tubercles on nape; tubercles on flanks conical, those of vertebral region somewhat flatter; dorsal body tubercles surrounded by 11–13 granular scales; dorsal tubercle rows at midbody 19–24; paravertebral tubercles between limb insertions 31–37, distinct vertebral row of tubercles absent; scales around midbody 118–130; ventral scales large; males with 25–30 precloacal pores in continuous series, females without precloacal pores (but pitted scales present); region posterior to vent covered by flat juxtaposed scales and greatly swollen, containing 1 or 2 enlarged tubercles on each side at level of vent; tail thick at base; light band on tail base in width of 9–12 granular scales and with 8–10 tubercles in transversal series.

Limbs relatively long and thin, covered dorsally with granular scales interspersed with several tubercles and ventrally with flat juxtaposed to subimbricate scales; dorsal granular scales grade into slightly flattened subimbricate scales on top of pes and manus; hind limbs larger than forelimbs; axillary pockets deep; subdigital lamellae wide, 10 or 11 under first finger, 18–21 under fourth finger, 11 or 12 under first toe, 21–25 under fourth toe; digits laterally compressed, increasing in length from first to fourth, fifth shorter than fourth (modified after Orlov et al. 2008; Nguyen 2011).

Coloration in life ( Fig. 3 )

Dorsal ground color of head, body and limbs signal brown (juveniles), grey-brown (young adults, SVL: 97.2–116 mm) and signal brown (adults), without small dark brown blotches (dark blotches present only on lower zone of flanks); dull white tubercles on dorsal body, limbs in juveniles, few orange tubercles on limbs in adults; iris orange or red brown; five bands on the dorsal ground, thin, immaculate without dark spots, slight bisque in juveniles, orange brown or yellow in adults, all edged anteriorly and posteriorly by thin dark brown bands, including one thin nuchal loop extending from posterior corners of eyes and posteriorly protracted (in V-shape), three body bands between limb insertions, and another one on tail base; ground color of tail dark brown, and signal brown at mottled tail base; 3–6 immaculate white caudal bands; ventral surfaces of head, body and limbs dull white and immaculate except for few dark brown spots on margin regions of gular and limbs.

Figures

Fig. 3. Goniurosaurus huuliensis Orlov, Ryabov, Nguyen, Nguyen & Ho, 2008. A. Juvenile (SVL=74.4 mm). B. Adult (SVL=97 mm). C. Adult (SVL=108 mm). D. Adult (SVL=116 mm). E. Adult (SVL=125 mm).F. Adult (SVL=132 mm).