Chimarra cakaudrovensis , Johanson, Kjell Arne & Oláh, János, 2012

Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, Zootaxa 3354, pp. 1-58: 38-40

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Chimarra cakaudrovensis

new species

Chimarra cakaudrovensis  , new species

Figs. 21View FIGURES 14 – 21, 111– 116View FIGURES 111 – 116

The genitalia of this species, particularly the gonopods, resemble those of C. helomyzida  , C. kimminsi  , C. cartwrighti  , C. lavensis  , C. devoensis  , C. manni  , C. devoensis  , C. tipulida  , and C. vanuensis  . It is easily distinguished from C. lavensis  , C. devoensis  and C. vanuensis  by the absence of prominent dorsad-oriented processes from the lateral lobes of tergum X; from C. kimminsi  , C. helomyzida  , C. tipulida  and C. cartwrighti  by the presence of 2 pairs of endothecal spines (not 1 pair); and from C. manni  by its strongly anterad produced segment IX. Tergum X somewhat resembles that of C. cartwrighti  and C. manni  , but the angle between the dorsal and ventral branches is much wider than in those 2 species, and the dorsal branches are more membranous than in C. manni  .

Male. Head and thorax dark brown, dorsal part of meso- and metathorax slightly darker brown than rest of thorax. Area between ocelli nearly black. Foreleg anterior claw as long as foreleg spur.

Wings ( Fig. 21View FIGURES 14 – 21). Forewings 5.0 mm (n= 1), brown. Forewings broad, ratio of length to breadth 3.2; R 1 slightly curved before crossvein r; radial sector slightly produced anterad immediately before discoidal cell; discoidal cell originating at mid-length of wing, about 3 x longer than wide; median cell as long as discoidal cell; crossvein r originating on R 2 + 3 immediately before fork I; fork I originating before crossvein s at distance equal to length of crossvein s; nygma located near base of fork II; fork III 1 / 5 th as long as wing; fork V slightly shorter than fork II; Cu 2 ending in wing margin well separated from A. Hind wings 4.0 mm (n= 1), brown; ratio of length to breadth 3.0; margin weakly incurved at arculus, where Cu 1 and Cu 2 fused with margin; fork I originating slightly before anterodistal corner of discoidal cell; fork III as long as discoidal cell and 1 / 7 th as long as wing; fork V as long as fork I; 1 A+ 2 A about 3 x as long as 1 A.

Male genitalia ( Figs. 111–116View FIGURES 111 – 116). Segment IX longer than high, nearly boomerang-shaped in lateral view; anterodorsal margins slightly produced anterad, ventrum strongly produced anterad into triangular plate in lateral view; ventral margins irregularly convex; anterodorsal margin deeply concave; each posterior margin convex; segment IX ventrally with with setae restricted to row before posterior margin ( Fig. 113View FIGURES 111 – 116). Dorsal part of segment IX nearly straight in lateral view, central part membranous in dorsal view; anterior margin invisible in dorsal view. In ventral view segment IX with almost parallel lateral, undulating margins, anterior and posterior margin shallowly concave; posterior margin without central projection. Tergum X short, divided into pair of lateral branches; in lateral view each lateral branch with small dorsal rounded lobe, and tube-shaped, nearly straight ventral branch with 2 apical sensillae. In dorsal and ventral view, ventral processes oriented mesad, crossing. Cerci located immediately below tergum X; wart-like in lateral and dorsal view, darkly pigmented; covered by long setae. Gonopods about as long as height of segment IX; nearly triangular in lateral view, basal 1 / 2 narrowing in lateral view; distal 1 / 2 nearly parallel-sided; each gonopod unbranched in lateral view; with well-developed dorsomesal lobe and ventromesal lobe of mesal process in ventral view. Anterodorsal margin of each gonopod nearly straight, smooth; posteroventral margin of basal 1 / 2 with strongly expanded setal bases, distal 1 / 2 with shorter setal bases; ventral margin strongly convex; apical margin armed with small megasetae. In ventral view, gonopods broad at base, forming rounded lobes with undulating margins; gonopods slightly curved mesad toward rounded apex. Mesal branch of each gonopod divided into well-developed ventromesal and dorsomesal lobe; ventromesal lobe edged, darkly pigmented, covered by minute setae; dorsomesal lobe strongly curved ventrad, with pointed apex. Phallic apparatus about 1.5 x longer than rest of genitalia, straight along its length, except slightly curved ventrad at posterior apex; phallotheca, in lateral view about 3 x thicker than posterior part; in ventral view about 2 x thicker than posterior part; apicoventral spine absent; phallotremal sclerite small, 2 -rayed, pointing posterad; 2 pairs nearly black, long, posterad and anterad directed endothecal spines present, about as long as diameter of narrowest part of phallotheca; endothecal spicules absent.

Female. Unknown.

Holotype male: TAVEUNI: Cakaudrove Prov., Devo Forest Reserve, Malaise trap, 3–10.i.2003, 16° 50 ’S, 179 ° 59 ’E [16.8333 °S, 179.9833 °E], 800 m, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc# 37] [ FNIC].

Paratypes: same data as holotype [loc# 37] [ NHRSAbout NHRS] — 11 males. Same data as holotype, except 10–16.i. 2003 [loc# 37] — 9 males [ BPBMAbout BPBM]. Cakadrove Prov., Davo Peak Radio Tower, Malaise trap in rainforest, 13–20.xii.2002, 16° 51 ’S, 179 ° 58 ’E [16.8500°S, 179.9667 °E], 1200 m, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc# 36] — 9 males [ BPBMAbout BPBM]. Same data, except 25.ix – 7.x. 2002 [loc# 36] — 1 male [ BPBMAbout BPBM]. Same data, except 10–17.x. 2002 [loc# 36] — 2 males [ BPBMAbout BPBM]. Same data, except 31.x – 21.xi. 2002 [loc# 36] — 1 male [ BPBMAbout BPBM]. VITI LEVU: Naitasiri Prov., Eteni, Navai,, Malaise trap, – 15.vii.2003, 17° 37 ’S, 177 ° 59 ’E [17.6167 °S, 177.9833 °E], 700 m, leg. E. Schlinger, M. Irwin & M. Tokota’a [loc#09] — 2 males [ NHRSAbout NHRS]. Vuda  Prov., Koroyanitu Pk., 1 km E Abaca Vlg., Malaise trap 1, 12– 19.xi. 2002, 17.667 °S, 177.55 °E, 800 m, leg. E. Schlinger & M. Tokota’a [loc#03] — 1 male [ NHRSAbout NHRS, DNAAbout DNA voucher IM 4]. Same data, except 19–26.xi. 2002 [loc#03] — 1 male [ BPBMAbout BPBM]. Same data, except 22.iv – 5.v. 2003 [loc#03] — 10 male [ BPBMAbout BPBM]. Naitasiri Prov., 3.2 km E Navai Vlg., Veilaselase Track, Malaise trap 2, 18– 25.x. 2004, 17.624 °S, 178.009 °E, 1020 m, leg. E. Schlinger & M. Tokota’a [loc# 15] — 1 male [ FNIC]. Rewa Prov., 3.8 km N Veisari, Waivudava Log Rd., Malaise trap, 25.iv – 25.v. 2003, 18.0792 °S, 178.3625 °E, 300 m, leg. E. Schlinger & M. Tokota’a [loc# 16] — 1 male [ NHRSAbout NHRS].

Etymology: Cakaudrovensis  , after Cakaudrove Province on Taveuni, the type locality of the species. Distribution: Taveuni and Viti Levu.


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