Chimarra veisarensis, Johanson, Kjell Arne & Oláh, János, 2012

Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, Zootaxa 3354, pp. 1-58 : 31-33

Chimarra veisarensis , new species

Figs. 17 , 90–94

Chimarra veisarensis has genitalia resembling those of C. schlingeri and C. nathani , from which it is easily separated by having forewings and hind wings each without a large pale spot in the center. In addition, tergum X forms 2 simple, lateral processes; and the dorsal branch of each gonopod is long, broad, straight, and directed posterad.

Male. Head and thorax pale brown, dorsal part of meso- and metathorax brown. Area between ocelli yellowish-brown. Foreleg anterior claw as long as foreleg spur.

Wings ( Fig. 17 ). Forewings 4.2 mm (n=1), brown. Forewings broad, ratio of length to breadth 3.3; R1 nearly straight before crossvein r; radial sector not produced anterad immediately before discoidal cell; discoidal cell originating at mid-length of wing, 3x longer than wide; median cell slightly shorter than discoidal cell; crossvein r fusing with basis of fork I; fork I originating before crossvein s at distance equal to length of crossvein s; nygma located near base of fork II; fork III 1 /7th as long as wing; fork V about as long as fork II; Cu2 ending in wing margin distantly from A. Hind wings 3.3 mm (n=1), brown; ratio of length to breadth 2.9; margin not incurved at arculus, where Cu1 and Cu2 fused with margin; fork III as long as discoidal cell and 1/8th as long as wing; fork V nearly as long as fork I; 1A+2A about 3x as long as 1A.

Male genitalia ( Figs. 90–94 ). Segment IX slightly taller than long; anterodorsal margins shallowly concave in lateral view; ventral margins irregularly convex; each posterior margin nearly straight, starting immediately below each cercus; ventral side of posterior 1/2 of segment IX densely covered by setae ( Fig. 92 ). In dorsal view with pointed anterior lobes; anterodorsal margin with wide, shallow incision in dorsal view, mesal margins with rounded process, seen also in lateral view. In ventral view segment IX with concave anterior margin and nearly straight posterior margin; posterior margin without central projection. Tergum X simple; tapering along its length, slightly curved ventrad along its length in lateral view ( Fig. 90 ); dorsal margin slightly convex, ventral margin slightly concave. In dorsal view tergum X divided basally into 2 lateral, nearly straight and diverging branches ( Fig. 91 ); each lateral branch tapering toward apex in dorsal and ventral view; pair of sensillae located subapically and apically on each lateral branch. Cerci short, wart-like in lateral view, located dorsally on segment IX, about at mid-height of basis of tergum X and oriented slightly posterodorsad in lateral view and oriented posterad in dorsal view; covered by long setae. Gonopods shorter than segment IX, ventral branch nearly quadrangular in lateral view; anterodorsal margin slightly concave, undulating; ventral margin nearly straight; dorsal branch forming thick, nearly straight process directed posterad, about as long as posterior margin of ventral branch of each gonopod, reaching as far posteriorly as tergum X. Ventral branch of each gonopod with straight posterior margin in lateral view; posterior margin with dense row of megasetae; in ventral view with sigmoid mesal margin. Mesal process situated centrally on gonopods, strongly sclerotized, slightly curved mesad. Phallic apparatus about 1.5x longer than rest of genitalia, slightly curved ventrad along its length; phallotheca, in lateral view nearly 3x thicker than posterior part, with anterior part about 2x thicker than posterior part; apicoventral spine absent; phallotremal sclerite complex, in ventral view with five posterior rays and1 anterior ray; pair of nearly black, short, posterad and anterad directed, subequally large endothecal spines present, slightly shorter than diameter of narrowest part of phallotheca.

Female. Unknown.

Holotype male: VITI LEVU: Naitasiri Prov., 3.3 km N Veisari, logging rd. to Waivudava, Malaise trap, 8–31.iii.2003, 18.0592°S, 178.367°E, 300 m, leg. M. Tokota’a [loc#20] [ FNIC].

Paratypes: Same data as holotype [loc#20] — 4 males [ NHRS]. Naitasiri Prov., 4 km WSW Colo-i-Suva Village, Mt. Nakobalevu, Malaise trap 1, 12.iv.2004, 18.057°S, 178.42°E, 300 m, leg E. Schlinger & M. Tokota’a [loc#12] — 1 male [ NHRS, DNA voucher IL 3]. Rewa Prov., 3.8 km N Veisari, Waivudava Log Rd., Malaise trap, 25.iv–25.v.2003, 18.0792°S, 178.3625°E, 300 m, leg. E. Schlinger & M. Tokota’a [loc#16] — 1 male [ BPBM].

Etymology: Ve i s are n s is, named after Veisari, near the type locality.

Distribution: Viti Levu.

Figures

FIGURES 14 – 21. Right wings, dorsal views. 14 — C. tokotaai, new species, holotype; 15 — C. vuda, new species, holotype; 16 — C. naitasirensis, new species, holotype; 17 — C. veisarensis, new species, holotype; 18 — C. fijiana, new species, holotype; 19 — C. abacensis, new species, holotype; 20 — C. manni Banks, non-type; 21 — C. cakaudrovensis, new species, holotype.

FIGURES 90 – 94. Chimarra veisarensis, new species, holotype. 90 — genitalia, left lateral view; 91 — genitalia, dorsal view; 92 — genitalia, ventral view; 93 — phallic apparatus, left lateral view; 94 — phallic apparatus, ventral view.