Chimarra naitasirensis, Johanson, Kjell Arne & Oláh, János, 2012

Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, Zootaxa 3354, pp. 1-58 : 30-31

Chimarra naitasirensis , new species

Figs. 16 , 85–89

Chimarra naitasirensis is similar to C. tokotaai and C. vuda in the presence of simple, dorsad-curved gonopods; a strongly enlarged phallotremal sclerite in the phallic apparatus; absence of large endothecal spines; and an endotheca with group of minute, strongly sclerotized spicules. Chimarra naitasirensis is distinguished from the other 2 species by having nearly drop-shaped gonopods in lateral view; a phallotremal sclerite that is about as long as the width of the posterior part of phallotheca; and by the presence of 3 groups of sclerotized spicules in the phallic endotheca.

Male. Body pale brown, dorsal part of meso- and metathorax brown. Large dark area between lateral and anterior ocelli. Foreleg anterior claw as long as foreleg spur.

Wings ( Fig. 16 ). Forewings 4.5 mm (n=1), brown; broad, ratio of length to breadth 3.3; R1 nearly straight before crossvein r; radial sector slightly produced anterad immediately before discoidal cell; discoidal cell originating at mid-length of wing, about 3x longer than wide; median cell slightly longer than discoidal cell; crossvein r situated basally of fork I by length of crossvein r; fork I originating before crossvein s at distance equal to 1/2 length of crossvein s; nygma located near base of fork II; fork III 1 /6th as long as wing; fork V longer than fork I, shorter than fork II; Cu2 situated close to A at wing margin. Hind wings 3.4 mm (n=1), brown; broad, ratio of length to breadth 3.0; margin slightly incurved at arculus, where Cu1 and Cu2 fused with margin; fork III slightly longer than discoidal cell and 1/8th as long as wing; fork V nearly as long as fork I; 1A+2A 4x longer than 1A.

Male genitalia ( Figs. 85–89 ). Segment IX about as long as high; parallelogram-shaped diagonal; each anterodorsal margin nearly straight in lateral view; each ventral margin convex, with strong incision at vertical apodeme; each posterior margin strongly produced posterad into sharply triangular lobe starting at top of cercus; ventral side of posterior 1/3rd of segment IX with setae ( Fig. 87 ). In dorsal view with narrow anterior lobes; anterodorsal margin forming deep U-shaped incision in dorsal view. In ventral view segment IX with shallowly triangular anterior margin and deeply U-shaped posterior margin; posterior margin without central projection. Tergum X simple; tapering apically in lateral view ( Fig. 85 ); orienting posteroventrad, nearly straight; divided into 2 lateral branches from base in dorsal and ventral view ( Figs. 86, 87 ); each branch uniformly narrowing posteriorly in dorsal and ventral view, apically pointed, without processes; sensillae invisible. Cerci large, broad, located immediately below dorsal margin of segment IX and tergum X and oriented posterodorsad in lateral view; sharply triangular in dorsal view; covered by long setae. Gonopodsunbranched, about as long as segment IX, uniformly curved dorsad, almost drop-shaped in lateral view; anterodorsal margin nearly straight; ventral margin uniformly convex; posteroventral corner not produced into tooth in lateral view; in ventral view forming broad plates with uniformly convex lateral margins; mesal margins diverging, except with large, 1/2-circular lobes at mid-length; tapering from 1/2-length into rounded, slightly mesad-curved apex. Phallic apparatus slightly longer than rest of genitalia: phallotheca, in lateral view, with anterior part slightly more than 3x thicker than posterior part; and ventral view anterior part about 2x wider than posterior part; apicoventral spine absent on phallotheca; strongly enlarged phallotremal sclerite in phallic apparatus forming complicated structure nearly 1/3rd as long as phallic apparatus; large endothecal spines absent; endotheca with 3 groups of minute, strongly sclerotized, well separated spicules; all groups densely packed in lateral view, more scattered in ventral view; anteriormost group located ventrally of posterior part of phallotremal sclerite; posterior 2 groups in longitudinal line with anterior group.

Female. Unknown.

Holotype male: VITI LEVU: Naitasiri Prov., Nakobalevu Mt., 24–29.x.2003, 18°03’S, 178°25’E [18.0500°S, 178.4167°E], 340 m, M. Irwin, E. Schlinger & M. Tokota’a [loc#10] [ FNIC].

Paratypes: S ame data as holotype [loc#10] — 2 males [ NHRS]. Vuda Prov., Koroyanitu Pk., 1 km E Abaca Vlg., Malaise trap 1, 22.iv–5.v.2003, 17.667°S, 177.55°E, 800 m, leg. E. Schlinger & M. Tokota’a [loc#03] — 2 males [ BPBM]. Vuda Prov., Koroyanitu Pk., 1 km E Abaca Vlg., Savuione Trail, Malaise trap, 7–12.x.2002, 17.667°S, 177.55°E, 800 m, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#03] — 1 male [ BPBM]. Same data, except 21.ix–7.x.2002 [loc#03] — 2 males [ FNIC].

Etymology: Naitasirensis , named after the Naitasiri Province, the type locality of the species. Distribution: Viti Levu.

Figures

FIGURES 14 – 21. Right wings, dorsal views. 14 — C. tokotaai, new species, holotype; 15 — C. vuda, new species, holotype; 16 — C. naitasirensis, new species, holotype; 17 — C. veisarensis, new species, holotype; 18 — C. fijiana, new species, holotype; 19 — C. abacensis, new species, holotype; 20 — C. manni Banks, non-type; 21 — C. cakaudrovensis, new species, holotype.

FIGURES 85 – 89. Chimarra naitasirensis, new species, holotype. 85 — genitalia, left lateral view; 86 — genitalia, dorsal view; 87 — genitalia, ventral view; 88 — phallic apparatus, left lateral view; 89 — phallic apparatus, ventral view.