Chimarra karoyanitensis , Johanson, Kjell Arne & Oláh, János, 2012

Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, Zootaxa 3354, pp. 1-58: 25-26

publication ID 10.5281/zenodo.210736

persistent identifier

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scientific name

Chimarra karoyanitensis

new species

Chimarra karoyanitensis  , new species

Figs. 13View FIGURES 6 – 13, 70– 74View FIGURES 70 – 74

Chimarra karoyanitensis  resembles C. schlingeri  , C. macuatensis  , and C. signata  in having gonopods each with a long, slender, ventrally hooked dorsal process. It is most similar to C. macuatensis  in the overall shapes of segment IX, tergum X, the gonopods, and the phallic apparatus. In lateral view, the gonopods of C. karoyanitensis  are broadest at mid-length, while in the other species the gonopods are broadest well beyond mid-length, as far out as the origin of the dorsal branch. It is distinguished from C. macuatensis  by the ventral branch of each gonopod posteriorly exceeding the dorsal branch, the longer ventral branches of tergum X, and the more rounded anterior lobes of segment IX in lateral view. It is distinguished from C. schlingeri  by the gonopods that are clearly longer than broad. It is separated from C. signata  by the straight posterior margins of segment IX in lateral view, and smaller dorsal branch of each gonopod relative to the ventral branch.

Male. Body yellowish brown, dorsal part of meso and metathorax brown. Large dark area between lateral and anterior ocelli. Foreleg anterior claw as long as foreleg spur.

Wings ( Fig. 13View FIGURES 6 – 13). Forewings 5.6 mm (n= 1), brown; ratio of length to breadth 3.3; R 1 nearly slightly undulating before crossvein r; radial sector slightly produced anterad immediately before discoidal cell; discoidal cell originating at mid-length of wing, 2 x longer than wide; median cell slightly shorter than discoidal cell; crossvein r situated at base of fork I; fork I originating before crossvein s at distance equal to 1 / 3 rd length of crossvein s; nygma located near base of fork II; fork III 1 / 5 th as long as wing; fork V slightly longer than fork I, shorter than fork II; Cu 2 close to A at wing margin. Hind wings 4.5 mm (n= 1), brown; broad, ratio of length to breadth 3.1; fork I with short footstalk, footstalk nearly as long as crossvein s; margin slightly incurved at arculus, where Cu 1 and Cu 2 fused with margin; fork III slightly longer than discoidal cell and 1 / 7 th as long as wing; fork V slightly shorter than fork I; 1 A+ 2 A 2 x longer than 1 A.

Male genitalia ( Figs. 70–74View FIGURES 70 – 74). Segment IX slightly shorter than high, nearly L-shaped in lateral view; anterior plates with large, nearly oval dorsal lobe, and small triangular lobe; posterior 1 / 2 of segment expanded dorsally; each anterodorsal margin deeply concave in lateral view; each ventral margin slightly convex, without incision at vertical apodeme; each posterior margin nearly straight, slightly undulating, angled 90 ° below cercus. In dorsal view segment IX with short, wide anterior lobes; anterodorsal margin forming shallow, wide, U-shaped incision without anterad-orienting processes on posteromesal margins. In ventral view segment IX nearly quadrangular, except narrow before anterior 1 / 3 rd; anterior margin widely concave; posterior margin straight, with short, wide central projection. Tergum X divided into dorsal and ventral branches, surrounding phallic apparatus. Tergum X with nearly straight dorsal margin in lateral view; dorsal branches rounded triangular, angled nearly 90 ° at posterior end; in dorsal view each lateral branch fused at base, 1 / 2 as wide as segment IX. Pair of ventral branches of tergum X smooth, slightly sigmoid an oriented posterad in lateral view; straight and diverging posterolaterad in dorsal view. Sensillae on tergum X not visible. Cerci located at dorsal margin of segment IX; straight, oriented posterodorsad in lateral view; covered by long setae. Gonopods slightly oval, broadest at mid-length, about as long as segment IX; dorsal margin convex, ventral margin nearly straight; 2 -branched; exceeding tergum X by length of cercus. Each dorsal branch slender, as long as individual cercus, strongly curved ventrad along its length, forming ventral hook; apex nearly as far out as apex of ventral branch; in dorsal view nearly straight, pointing posteromesad; mesal megasetae absent. Ventral branch of each gonopod nearly hyperbolic in lateral view, with undulating posterior and ventral margin; each with mesal margin strongly undulating in ventral view. Mesal branches absent. Phallic apparatus slightly longer than rest of genitalia, nearly straight; phallotheca, in lateral view, with anterior part 3 x thicker than posterior part; in ventral view anterior part about 2 x wider than posterior part; apicoventral spine absent on phallotheca; phallotremal sclerite not observed; 2 long, nearly black endothecal spines present.

Female. Unknown.

Holotype male: VITI LEVU: Vuda  Prov., Koroyanitu Natural Heritage Park, Savuione Trail, Malaise trap, 12–19.x.2002, 17° 40 ’S, 177 ° 33 ’E [17.3333 °S, 177.55 °E], 450 m, M. Irwin, E. Schlinger & M. Tokota’a [loc#04] [ FNIC].

Paratypes: Same data as holotype, except: 21.ix – 7.x. 2002 [loc#04] — 2 males [ NHRS]. Viti Levu, Vuda  Prov., Koroyanitu Pk., 1 km E Abaca Vlg., Malaise trap 1, 7– 12.x. 2002, 17.667 °S, 177.55 °E, 800 m, leg. E. Schlinger & M. Tokota’a [loc#03] — 2 males [ BPBM]. Same data, except: 19–26.x. 2002 [loc#03] — 1 male [ NHRS]. Same data, except: 26.x – 5.xi. 2002 [loc#03] — 2 males [ BPBM]. Same data, except: 26.xi – 3.xii. 2002 [loc#03] — 3 males [ NHRS]. Same data, except: 10–17.xii. 2002 [loc#03] — 1 male [ FNIC].

Etymology: Karoyanitensis  , named after Koroyanitu Natural Heritage Park, the type locality of the species.

Distribution: Viti Levu.


Swedish Museum of Natural History, Entomology Collections


Bishop Museum