Ptychoderes brevis Jordan , 1894

Mermudes, José Ricardo M. & Napp, Dilma Solange, 2006, Revision and cladistic analysis of the genus: Ptychoderes Schoenherr, 1823 (Coleoptera, Anthribidae, Anthribinae, Ptychoderini), Zootaxa 1182 (1), pp. 1-130 : 26-28

Ptychoderes brevis Jordan, 1894

Figs. 209 , 230 , 242 , 263

Ptychoderes brevis Jordan, 1894: 592 ; Bovie, 1906: 219 (cat.); Wolfrum, 1929: 6 (cat.); Blackwelder, 1947: 765 (cat.); Rheinheimer, 2004: 10 (cat.).

Female. Vestiture ( Fig. 209 ): pale dorsal vitta with dense whitish scales, intermingled with olivaceous­green scales, decumbent and denser at sides of pronotum, elytral interstriae 1, 2 and 4 and apical declivity. Pygidium with predominance of olivaceous­green scales. Ventral body surface and legs with appressed, dense, yellowish­white scales; sides of prosternum, below lateral carina, with a vitta of scales concolorous with those of pronotum ( Fig. 230 ). Sides of mesosternum with a wide, glabrous vitta. Mesepimera, mesepisterna, metepisterna, sides of both metasternum and ventrites with shallow setigerous punctures, each provided with a pale, yellowish scale. Outer surface of meso­ and metafemora with two spots of blackish­brown scales.

Rostrum at least 1.2 times longer than wide across base. Distance between scrobe and eyes about half maximum width of eye. Sides looking tricarinate due to the careniform upper margin of scrobe; carinae long and thick fused with the scrobal margin; with a transverse carina between the margin of scrobe and lower carina. Scrobes careniform at dorsal and basal margins; extension deep, narrow with a deep, distal pit, slightly wider than extension. Mentum finely, sparsely punctate; strongly impressed at sides, shallowly grooved at apical margin. Gula with moderately coarse, shallow, sparse punctures. Frons with prominent carinae.

Antennae short, reaching antebasal carina of prothorax. Segments III–IV slightly longer than II; V slightly longer than VI–VIII, subequal; IX one third longer than VIII; X the shortest; IX–X weakly expanded at apex; XI one third longer than X (II = 0.56–0.65; III = 0.70–0.72; IV = 0.72–0.75; V = 0.60–0.65; VI = 0.52–0.55; VII = 0.52–0.55; VIII = 0.48–0.50; IX = 0.75–0.76; X = 0.48–0.55; XI = 0.80).

Prothorax ( Figs. 209 , 242 ) slightly longer than wide; sides converging anteriorly from antebasal carina. Pronotal depression deep, with a almost rounded, weakly prominent, median tubercle, not reaching lateral margins of depression. Pronotum evidently rugose throughout, the rugae prominent and long. Antebasal carina interrupted at middle (almost twice width of carina), oblique towards lateral carina. Lateral carina straight ( Fig. 230 ). Secondary carina sinuous, narrowly interrupted at middle, almost reaching lateral carina. Laterobasal carina reaching posterior margin of prothorax. Prosternum raised, with coarse, deep dense punctures; antecoxal furrow straight, deep and entire. Mesosternum with coarse, shallow, sparse punctures; near the intercoxal process with two median, almost rounded, contiguous, deep depressions and coarse punctures at sides. Intercoxal process of mesosternum evidently impressed at apex, swollen at lateral and apical margins, with lateroapical projections. Elytra 1.2–1.9 times longer than wide across humeri, somewhat convex, transversely depressed before middle; basal gibbosities prominent; surface rugose from base to behind middle, more so at base and near suture. Sides subparallel at basal half, slightly wider than humeri behind middle. Anteapical tubercles absent, interstriae not swollen. Outer apical glabrous area of profemora with a single deep puncture.

Abdomen. Ventrite I with scarce, fine punctures, partially obscured by vestiture; intercoxal process depressed, groove line shaped. Ventrite V one third longer than IV, with lateroapical projections somewhat evident, not acuminated; apex emarginate. Pygidium slightly longer than wide, subconvex, not impressed medially at apex; with a short groove to fit elytra; apex truncate. Ovipositor (apex exposed) with a toothed plate similar to that of P. crustatus .

Measurements in mm, female. TL = 14.3–14.5; RL = 2.0–2.1; RAW = 1.7–1.9; RBW = 1.5–1.7; HW = 2.3–2.5; DSO = 0.7–0.7; MLO = 0.7–0.7; PL = 4.2–4.3; PW = 4.0–4.2; E L = 6.0–9.0; EW = 4.7–4.8.

Male. Unknown.

Type material

Female holotype ( BMNH) examined, labelled as follows: 1) Type [rounded, white with red border, printed]; 2) Ptychoderes brevis , ♀, Type: Jordan [white, manuscript]; 3) Pará, N. Brazil [white, printed]; 4) Karl Jordan coll., B.M. 1940–109 [white, printed]. Type locality: Brazil (Pará) .

Remarks

Ptychoderes brevis belongs to the P. crustatus + group, being the sister group of P.jekeli Mermudes and Napp, 2004 by two synapomorphies: pronotal depression with a median, somewhat rounded, weakly prominent tubercle (20 5), and glabrous area of profemora with a single coarse puncture (36 2). P. brevis differs from P. jekeli , and also from all other species of Ptychoderes , by the pygidium not impressed medially near apex, and ventrite V with projected, not acuminated, lateroapical angles, both autapomorphies of P. brevis .

Ptychoderes brevis is similar to P. callosus Jekel, 1855 by the dorsal vestiture with olivaceous­green scales, differing by: 1) prosternum raised with coarse, deep, dense punctures; 2) prosternal furrow straight, deep and entire and 3) elytra with prominent basal gibbosities, lacking both anteapical tubercles and swollen interstriae. In P. callosus , the prosternum is weakly raised, with moderately coarse, shallow, sparse punctures; the prosternal furrow is shallow, strongly curved in front of procoxae, and with a subtriangular prosternal projection at middle not interrupting the furrow; the elytra have weakly prominent basal gibbosities, anteapical tubercles and interstriae 3–9 swollen.

Distribution ( Fig. 263 ) COLOMBIA and BRAZIL (Pará, Mato Grosso).

Material examined

COLOMBIA. Putumayo: Nariño (Villa Amazonica), 1 F (compared with holotype), 1.X– XI .1995, Mracek col. ( CPMT) . BRAZIL. Mato Grosso: Sinop, 1 F, Isnard Rubim leg. ( MZSP) .

Figures

FIGURES 208–214. 208, Ptychoderes crustatus Frieser, 1988, male; 209, Ptychoderes brevis Jordan, 1894, female holotype; 210, Ptychoderes jekeli Mermudes & Napp, 2004, female holotype; 211, Ptychoderes longicollis Jordan, 1894, male. 212–214, Ptychoderes jordani Frieser, 1959. 212, male holotype; 213, female paratype; 214, pale form, male.

FIGURES 228–238. 228, Ptychoderes bivittatus Jordan, 1894, male; 229, P. nebulosus (Olivier, 1795), male; 230, P. brevis Jordan, 1894, female holotype. 231–234 detail of elytral apex. 231, Unanthribus maximus; 232, Ptychoderes mixtus; 233, Ptychoderes elongatus; 234, Ptychoderes callosus. 235, pygidium and ventrite V, Ptychoderes nebulosus. 236–238, ventrite I, male. 236, Ptychoderes mixtus; 237, Ptychoderes rugicollis; 238, Ptychoderes nebulosus.

FIGURES 239–250. Pronotum. 239, Tribotropis prasinata; 240, Unanthribus grandis; 241, Ptychoderes crustatus; 242, P. brevis, female holotype; 243, P. longicollis, holotype; 244, P. jordani, male holotype; 245, P. magnus, male holotype; 246, P. depressus; 247, P. virgatus; 248, P. bivittatus, male syntype; 249, P.rugicollis; 250, P. elongatus.

FIGURES 263–264. 263. Geographical distribution of P. longicollis (black circles), P. crustatus (white triangles), P. brevis (black squares), P. jekeli (black triangles). 264. Geographical distribution of P. jordani (black squares), P.depressus (black triangles), P. virgatus (white circles).