Atopida suturalis ( White, 1846 )

Kiałka, Agata & Ruta, Rafał, 2022, Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae), Zootaxa 5174 (4), pp. 401-443 : 429-432

Atopida suturalis ( White, 1846)

( Figs 2G–H , 3F , 4 , 5S –T, 6A , 7A–B , 8K–L , 10J–K , 15C , 17B )

Ptinus suturalis White, 1846: 8 .

Atopida suturalis: Sharp 1884: 299 ; Pic 1914: 16 (catalogue); Macfarlane et al. 2011: 420 (catalogue); Kiałka & Ruta 2017: 26 (catalogue).

Atopida brouni Sharp, 1878: 47 .

Atopida brouni: Broun 1880: 312 (reprinted excerpt of Sharp 1878); Sharp 1884: 299 (synonym of Atopida suturalis (White)) ; Pic 1914: 16 (catalogue); Macfarlane et al. 2011: 420 (catalogue); Kiałka & Ruta 2017: 27 (catalogue).

Atopida lawsoni Sharp, 1878: 47 syn. nov.

Atopida lawsoni: Broun 1880: 312 (reprinted excerpt of Sharp 1878); Pic 1914: 16 (catalogue, as a synonym of A. suturalis ); Macfarlane et al. 2011: 420 (catalogue); Kiałka & Ruta 2017: 24 (catalogue).

Type material. Holotype of Ptinus suturalis (NHM) : pinned into card, “Type” [round label with red border], “ Ptinus suturalis White \ Zool. Ereb & Terror. \ Leptosp. flowers \ Hooker and Smith” [handwritten]. Holotype of A. lawsoni , female ( NHM): card-mounted, “ Atopida Lawsoni \ Type D.S. \ Auckland Lawson” [written at base of card in Sharp’s hand], “Type \ H. T.” [round label with red border], “ Auckland, [red underline] \ New Zealand.”, “Sharp Coll. \ 1905-313.” Syntypes of A. brouni , 2 exx. ( NHM): card-mounted, “ Atopida brouni \ Type D.S. \ Tairua Broun” [written at base of card in Sharp’s hand], “Type \ H. T.” [round label with red border], “Tairua, [red underline] \ New Zealand. \ Broun.”, “Sharp Coll. \ 1905-313.” Syntype, 1 ex. ( NHM): card-mounted, “ Atopida brouni \ Ind. typ. \ D.S. \ Tairua Broun” [written at base of card in Sharp’s hand], “Tairua, [red underline] \ New Zealand. \ Broun.”, “Sharp Coll. \ 1905-313.”, “= suturalis White ” [handwritten] .

Other material examined. 3 males ( AMNZ): “ Wayby Gorge \ N. Auck. \ 28.12.26 [on the underside of the cardboard with the beetle]”, “ C.E. Clarke \ Collection ”, “ Atopida \ suturalis \ White \ = brouni Shp. \ Id. by J. Armstrong \ comp. Cot. brouni” ; 1 ex. ( CMNC): “ NEW ZEALAND \ North Is. Huia \ 17-27.XI.1981 \ H & A Howden”; 7 exx. ( DBET): “New Zealand AK 1: \ Waikatere Range, \ Waiatarua W, -36.9372, \ 174.5575, 260 m, beating \ in wet forest, 26.11.2016, \ leg. R. Ruta ”; 14 exx. ( DBET): “ New Zealand ND 24: \ Waipoua Forest , Yakas \ Track, 7.12. 2016, 300 m \ beating, leg. R. Ruta ” ; 1 ex. ( DBET): “New Zealand ND 25: \ Waipoua Forest, Yakas \ Track , 300 m, 7.12.2016, \ litter sample, leg. R. Ruta ”; 2 exx. ( DBET): “ New Zealand ND 29: \ Toatoa Track, - 35.67, \ 173.56, 270 m, beating, \ 7.12.2016, leg. R. Ruta ”; 17 exx. ( CMNC): “ N. Zealand, ND \ Waipoua Kauri Forest \ Dec. 11&12, 1983 \ L. Masner ”; 4 exx. ( CMNC): “ NEW ZEALAND: N. Is. \ Waipoua S. Forest \ 20- 23.XI.1981 \ H & A Howden”; 2 exx. ( NZAC): “Waipoua \ S. F. Oct. 67 \ Auckland ”, “at light \ camp”, “ J.C. Watt ” ; 1 ex. ( NZAC): “ Waipoua S. F. \ Auckland \ 12 Oct. 67”, “at light \ J.C. Watt ” ; 1 ex. ( LUNZ): “ Waipoua For. \ 7.12.61 \ G. Kuschel ” ; 1 ex. ( LUNZ): “ Waipoua S. F. \ 16.X.67”, “to m/v light \ J.S. Dugdale ” ; 1 ex. ( LUNZ): “ NEW ZEALAND ND \ Waipoua SF \ 4 Feb 1975 \ sweeping”; 2 exx. ( NZAC): “Waipoua Beach \ 18 Oct. 67 \ Auckland ”, “at light”, “ J.C. Watt ” ; 1 ex. ( NZAC): “ NEW ZEALAND ND \ Waipoua Forest \ 1-2 Dec 1989 \ D.J. Bickel \ Pan traps ” ; 1 ex. ( NZAC): “ NEW ZEALAND ND \ Waipoua SF \ Waipoua River \ 31 Oct 1985 \ R.C. Craw” ; 1 female ( NZAC): “ NEW ZEALAND ND \ Waipoua SF \ 26 Nov 1980 \ G. Kuschel” ; 1 male ( NZAC): “ Waipoua \ S.F. Oct. 67 \ Auckland ”, “m/v. \ light.”, “ J.S. Dugdale ” ; 1 male ( NZAC): “988”, “ New Zealand ND \ Waipoua Forest \ 25.11.94”, “ Ex John Nunn \ Collection \ Donated 2015” ; 2 males ( NZAC): “ NEW ZEALAND ND \ Waipoua SF \ 26 Nov 1980 \ G. Kuschel” ; 1 male ( NZAC): “ NEW ZEALAND, ND \ Waipoua SF \ Lookout \ 29 Oct 1980 \ G. Kuschel” ; 1 male ( NZAC): “ NEW ZEALAND, ND \ Waipoua SF \ Lookout \ 29 Oct 1980 \ G. Kuschel” ; 1 male ( NZAC): “ Kawerua \ Beach ”, “19.x.67 \ J.S. Dugdale ”, “Waipoua S.F.” ; 1 ex. ( NHM): “Hunua G. \ nr Papakura \ 13-12-36 \ A. Richardson ”, “ NEW ZEALAND \ C.E. Clarke \ Collection \ B. M. 1957-24.”; 2 exx. ( LUNZ) , 1 female ( NZAC): “ Hunua Falls \ J.C. Watt \ 2 Dec 61 AK”, “ Dacrydium \ cupressinum \ Podocarpus \ totara”; 3 exx. ( NZAC): “ Hunua Ra. \ 2.12.61 \ G. Kuschel ” ; 2 males ( AMNZ): “ Hunua \ Gorge \ 13-12-36 \ A. Richardson [on the underside of the cardboard with the beetle]”, “HUNUA G. \ nr. Papakura \ 13-12.36 \ A. Richardson ”, “ Manukau Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM”, “ C.E. Clarke \ Collection ”, “ C. ocularis ” ; 1 male ( NHM): “ Waitakere Ra \ Destruction \ Gully \ 2.xi.1985 \ C & J Lyal”, “General \ beating”, “ NEW ZEALAND \ AK (CL 94) \ BM 1986-274” ; 2 females ( NZAC): “ NEW ZEALAND \ AK (CL 94) \ BM 1986-274”, “ Waitakere Ra \ Destruction \ Gully \ 2.xi.1985 \ C & J Lyal”, “General \ beating” ; 1 male ( AMNZ): “31.12.47 \ Waitakere \ Ras. C.T.”, “ Waitemata Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM”, “ C.E. Clarke \ Collection ”, “ Atopida \ sp.” ; 1 female ( NZAC): “ Waitakeri \ 30-12-15”, “560”, “T. Broun \ Collection ”, “ A.E. Brookes \ Collection ” : 1 male ( NZAC): “ Waitakeri \ 31-12-14”, “561”, “T. Broun \ Collection ” ; 1 female ( NZAC): “ NEW ZEALAND AK \ Waitakere Ra \ Fairy Falls Trk \ 4 Dec 1979 ”, “C.F. Butcher and \ M.F. Tocker \ On flowers of \ Leptospermum \ scoparium” ; 1 female ( NZAC): “ NEW ZEALAND AK \ Waitakere Range \ Cutty Grass Trk \ 2 Jan 1984 \ R.C.&J.P.N. Craw” ; 1 male ( NZAC): “WAITAKERES \ AUCKLAND \ 9.DEC.1953 \ A.E. BROOKES.” ; 1 male ( NZAC): “ Anawhata Rd. \ Waitakere \ 1Jan1945 \ M.W. Carter ”; 2 exx. ( NHM): “ NEW ZEALAND \ ND (CL 55) \ BM 1986-274”, “Omapere \ 30.X.1985 \ C & J Lyal”; 3 exx. ( NZAC): “Pandora”, “ Spirits Bay \ Northland \ Nov 67 \ J.I. Townsend ” ; 1 ex. ( NZAC): “ NEW ZEALAND ND \ Unuwhao, 270m \ 25 Nov 198X \ G. Kuschel ”, “sifted litter \ and decayed \ wood 82/125”; 2 exx. ( NZAC): “Uauwhao 1000’ \ Spirits Bay ”, “18.1.66 \ A. K. Walker ”, “Rhopalostylis \ sapida” ; 1 ex. ( NZAC): “ Unuwhao 243’ \ Spirits Bay ”, “18.I.66 \ A. K. Walker ” ; 10 males, 2 females ( NZAC): “ NEW ZEALAND ND \ Unuwhao , 270m \ 25 Nov 1982 \ G. Kuschel”, “Sifted litter \ and decayed \ wood 82/125”, “Duplicate \ specimens \ in alcohol” ; 2 males ( NZAC): “ NEW ZEALAND ND \ Unuwhao , 270m \ 26 Nov 1982 \ G. Kuschel” ; 1 ex. ( NZAC): “ NEW ZEALAND CL \ Little Barrier I \ 183m 29 Feb 1976 \ A.K. Walker \ sweeping” ; 1 ex. ( NZAC): “ NEW ZEALAND AK \ Riverhead State \ forest Reserve \ 11 Nov 1980 \ M.F. Tocker”, “on \ Myrsine \ australis” ; 1 ex. ( NZAC): “ NEW ZEALAND \ TV \ 28 Dec 1997 \ P A Maddison”; 5 exx. ( NZAC): “ NEW ZEALAND ND \ Paihia Opua SF \ 22 Jan 1981 \ G Kuschel” ; 1 ex. ( NZAC): “ NEW ZEALAND, TK \ Piopio, Mangaotaki \ Bush Walk \ 8 Dec 2000, G. Hall \ R. Hoare, T. Sirey”, “under logs \ Podocarp forest \ 38°28’S, 174°51’E ” ; 1 ex. ( NZAC): “ NEW ZEALAND ND \ Warawara camp \ 15 December 2007 \ T R Buckley D Seldon \ R Hoare”, “to MVL \ TB107 \ 35 22.167 S \ 173 17.012 E ” ; 1 ex. ( NZAC): “ NEW ZEALAND CL \ Rd E of Tapu \ 450 m Summit \ 3 Dec 1989 \ D. J. Bickel ”; 2 exx. ( NZAC): “Swanson \ X Totara \ 18-12-55 J.C. Watt \ 439B”; 3 exx. ( NZAC): “Swanson \ X Karamu \ 18- 12-55 J.C. Watt \ 439B” ; 1 ex. ( NZAC): “ Swanson \ X Olearia \ 18-12-55 J.C. Watt \ 439B” ; 1 ex. ( NZAC): “Swanson \ X Tanakaha \ 18-12-55 J.C. Watt \ 439B”; 5 exx. ( NAZC): “Swanson \ X Kanuka \ 18-12-55 J.C. Watt \ 439B”; 2 exx. ( NZAC): “ Swanson \ Auckland \ 13 Dec 58”, “ J.C. Watt ” ; 2 males ( NZAC): “ Swanson \ Auckland \ 13 Dec 58”, “ J.C. Watt ” ; 1 female ( NZAC): “ Ex light trap \ Titirangi \ NOV/51 \ C.R. Thomas. ”; 5 exx. ( NZAC): “ Ex light trap \ Titirangi \ NOV/52 \ C.R. Thomas. ” ; 1 female ( NZAC): “ Ex light trap \ Titirangi \ DEC/52 \ C.R. Thomas. ”; 2 exx. ( NZAC): “ Ex light trap \ Titirangi \ DEC/53 \ C.R. Thomas. ” ; 1 male ( NZAC): “ Titirangi \ 28 Nov 1942 \ M.W. Carter” ; 1 ex. ( NZAC): “ NEW ZEALAND AK \ Auckland \ Titirangi \ 15 Oct 1978 \ P.A. Maddison”, “At \ light” ; 2 females ( NZAC): “ Titirangi ”, “P.E.B. 27-12-49”, “ A.E. Brookes \ Collection ” ; 2 females ( NZAC): “ NEW ZEALAND AK \ Titirangi \ Minnehaha Res \ 9 Dec 1980 \ P.A. Maddison” ; 1 female ( NZAC): “ NEW ZEALAND AK \ Titirangi \ Minnehaha Res \ 9 Dec 1980 \ P.A. Maddison”, “Beating \ streamside \ shrubs” ; 1 female ( NZAC): “ NEW ZEALAND AK \ Auckland \ Titirangi \ 10 Dec 1978 \ P.A. Maddison”, “In house” ; 2 males ( NZAC): “ NEW ZEALAND AK \ Auckland \ Titirangi \ 15 Dec 1978 \ P.A. Maddison ”, “At light”; 2 exx. ( NZAC): “ Waimatenui \ Northland \ North Is. ”, “ Collector \ 10.12.1951 \ A.E. Brookes ” ; 1 female ( NZAC): “ Waimatenui \ V.1715.10.12.69 \ Brooks ”, “ A.E. Brookes \ Collection ” ; 1 female ( NHM): “ Paparoa ”, “ Atopida \ lawsoni”, “ New Zealand. \ Broun Coll. \ Brit.Mus. \ 1922-482.”, “lawsoni \ Broun coll.” ; 1 male, 1 female ( NZAC): “980”, “ New Zealand ND \ Radar Bush \ Te Paki \ 4.12.95”, “ In forest \ litter”, “ Ex John Nunn \ Collection \ Donated 2015” ; 1 female ( NZAC): “ NEW ZEALAND ND \ Te Paki Trig \ 23 Nov 1982 \ G.Kuschel”, “Sifted litter \ and decayed \ wood 82/126” ; 2 males ( NZAC): “ NEW ZEALAND ND \ Te Paki Trig \ 25 Nov 1982 \ G. Kuschel”, “Sifted litter \ and decayed \ wood 82/117” ; 1 female ( NZAC): “ New Zealand WN \ Pakuratahi Forks \ Katioke \ 7-10 Feb 92”, “ Ex John Nunn \ Collection \ Donated 2015” ; 1 female ( AMNZ): “ NEW ZEALAND, ND \ Waima R catchment \ Mangatete Stm \ 15.x.1985, R.F. Gilbert \ Swept foliage by stream”, “AMNZ126025 \ AUCKLAND \ MUZEUM \ NEW ZEALAND ” ; 1 male ( NZAC): “ NEW ZEALAND ND \ Long Louis \ Waima Forest \ 13 Feb 2008 \ R Leschen B Michaux”, “beating \ RL13330 \ 35°32.222’S 173°27.238’E ” ; 1 male ( AMNZ): “ex. Folliage \ Walkers Bsh \ Nov. 1966. MBP.”, “ Waitemata Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM”, “ M.B. Paterson \ Collection”, “AMNZ126073 \ AUCKLAND \ MUZEUM \ NEW ZEALAND ” ; 1 male ( NZAC): “NORTHCOVE”, “561”, “T. Broun \ Collection ” ; 1 male ( NZAC): “ New Zealand ND \ Omahuta SF \ Nov-94”, “ In leaf \ litter”, “ Ex John Nunn \ Collection \ Donated 2015” ; 1 male ( NZAC): “ NEW ZEALAND, ND \ Pukenui F \ 14 Sep 1998 \ G. Hall \ under logs” ; 1 male ( NZAC): “ NEW ZEALAND, AK \ Hillsborough \ 2001 \ D.S. Seldon \ Pittosporum tree” ; 1 male ( NZAC): “ NEW ZEALAND WN \ Stakas Valley \ 23 Feb 1975 \ B.A. Hallaway” ; 1 male ( AMNZ): “ Albany \ Hobson Block \ 11.XI.1982 \ R.F. Gilbert” ; 1 male ( AMNZ): “ NEW ZEALAND AK \ Atuanui \ Mt Auckland \ pitfall F 25 \ Jan 2002, A. Warren” ; 2 males ( AMNZ): “ NEW ZEALAND, ND \ E of Puketi SF \ Mangirangina Scientific \ Reserve , 12.xi.1985 \ R.F. Gilbert”, “Swept entrance to \ walkway” ; 1 male ( AMNZ): “swept understorey \ of high forest \ Waikaraka Valley \ S. of Puketi Forest \ 13.XI.1985 \ R.F. Gilbert \ NZMSI NII 273527 \ 35°16’ 173°48’” ; 1 male ( AMNZ): “swept understorey \ Waikaraka Strm.Vy. \ S. of Puketi Forest \ 14.XI.1985 ” ; 1 male ( AMNZ): “ NEW ZEALAND AK \ Mataitai Forest S.A. \ pitfall D 25 \ Jan 2002 \ A. Warren” ; 2 males ( AMNZ): “MANGAMUKA G. \ G.Z. 64 \ R.A. CUMBER” ; 1 male ( AMNZ): “ Mangamuka ”, “ Mangonui Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM”, “ C.E. Clarke \ Collection ” .

Diagnosis. Body relatively large (TL 4.5–5.3 mm), light brown to brown, head and pronotum usually a tone darker than elytra, setation of elytra adherent. Externally resembling A. castanea but pronotum of females with sparse punctation in basal half, elytra of females often yellowish and contrasting with dark brown pronotum and head. Penis moderately elongated, trigonium relatively wide, with rounded apex and parallel sides, as long as parameroids; pala slightly shorter than parameroids. Bursal sclerite long, with hooked processes in central portion.

Redescription. Male. Body elongated, moderate to large (TL 4.5–5.3 mm), moderately convex. Body brown, elytra often lighter than head and pronotum. Punctation on head and pronotum granulate, strong and dense, punctures doughnut-shaped, touching each other, punctation on elytra simple, punctures separated by 0.5–1.0× diameter of a puncture. Body covered with relatively short setae, semi-erect on head and pronotum, and adherent on elytra. Supraantennal ridges well-marked, elevated. Pronotum widest in anterior 1/3, sides curved, anterolateral angles slightly produced, posterolateral angles slightly obtuse. Elytra without carinae, humeri well-marked. Ventrite 5 rounded, with triangularly rounded apex. Penis (L 0.68 mm, W 0.29 mm) moderately elongated, trigonium narrow, rectangular, with rounded apex and parallel sides, as long as or slightly longer than parameroids; parameroids elongated, apices pointing outwards, covered with numerous punctures, inner edge slightly irregular; pala triangular, about the same length as parameroids. Tegmen (L 0.59 mm, W 0.33 mm) moderately wide, parameres slightly tapered, with scarce punctures and very few delicate setae on the inner edge.

Female. Body moderate to large (TL 4.5–5.2 mm), elongate, moderately convex, macropterous. Body brown, head and pronotum darker, elytra contrasting yellowish-brown, legs and antenna lighter. Supraantennal ridges wellmarked, slightly elevated triangularly above bases of antennae. Pronotum with peculiar punctation—punctures in basal portion sparse, punctures separated by 2.0× diameter of a puncture, interspaces glabrous, remaining areas of pronotum covered with coarse, granulate punctures. Anterolateral angles of pronotum more produced than in males, triangular. Elytra covered with very short, adherent setae, distinctly shorter than in males, not reaching next puncture, punctation of elytra coarser than in males. Ventrite 5 rounded, with triangularly emarginated apex. Lateral portions of ventrites 3 and 4 with small, oval setal patches. Bursal sclerite long (L 0.40 mm), anterior and posterior portion with two subparallel rod-like structures, central portion subtriangular, with well visible hooked projections on sides.

Measurements and ratios. Males (n = 10) TL 4.50–5.30 mm (4.98 mm), PL 0.75–0.90 mm (0.86 mm), PW 1.25–1.45 mm (1.39 mm), EL 3.75–4.50 mm (4.16 mm), EW 1.95–2.40 mm (2.15 mm), TL/EW 2.21–2.38 (2.32), PW/PL 1.53–1.73 (1.62), EL/EW 1.86–2.00 (1.93), EL/PL 4.41–5.29 (4.87), EW/PW 1.48–1.68 (1.55). Females (n = 10) TL 4.50–5.20 mm (4.76 mm), PL 0.75–0.95 mm (0.84 mm), PW 1.35–1.60 mm (1.48 mm), EL 3.80–4.20 mm (3.98 mm), EW 1.95–2.25 mm (2.11 mm), TL/EW 2.14–2.36 (2.11), PW/PL 1.67–1.93 (1.76), EL/EW 1.81–1.98 (1.89), EL/PL 4.42–5.07 (4.74), EW/PW 1.34C1.50 (1.43).

Distribution. Widely distributed in North Island, common in Northland and Auckland areas, less common in Coromandel, Waikato, and Wellington areas ( Fig. 18J ).

Figures

FIGURE 2. Atopida, dorsal view. A) A. paparoa sp. nov., female; B) A. paringa sp. nov., male; C) A. proba, male; D) same, female; E) A. suffusa, male; F) same, female; G) A. suturalis, male; H) same, female; I) A. tuhua, male; J) same, female; K) A. villosa, male; L) same, female; M) A. waipoua sp. nov., male; N) A. walkerae sp. nov., male; O) A. westlandensis sp. nov., male; P) same, female. Scale bars = 2.0 mm.

FIGURE 3. Atopida, morphological details. A) A. castanea, head, anterior view; B) same, ventral portion of prosternum and mesoventrite; C) A. insularis sp. nov., ventral portion of head, prosternum and mesoventrite; D) same, head and pronotum, dorsal view; E) A. lescheni sp. nov., male, head, projection of supraantennal ridge marked with arrow; F) A. suturalis, female, head, projection of supraantennal ridge marked with arrow. Abbreviations: mp—mesoventral process, pp—prosternal process, sgr—subgenal ridge, soc—subocular carina.

FIGURE 4. Atopida suturalis, mouthparts. A) mandible, B) maxilla, C) labrum, D) labium. Scale bar = 0.2 mm.

FIGURE 5. Atopida, pronotum. A) A. castanea, male; B) same, female; C) A. hirta, male; D) same, female; E) A. impressa, male; F) same, female; G) A. insularis sp. nov., male; H) same, female; I) A. lescheni sp. nov., male; J) A. montana, male; K) same, female; L) A. paparoa sp. nov., male; M) same, female; N) A. paringa sp. nov., male; O) A. proba, male; P) same, female; Q) A. suffusa, male; R) same, female; S) A. suturalis, male; T) same, female; U) A. tuhua sp. nov., male; V) same, female; W) A. villosa, male; X) same, female; Y) A. waipoua sp. nov., male; Z) A. walkerae sp. nov., male; AA) A. westlandensis sp. nov., male; BB) same, female. Not to the same scale.

FIGURE 6. Atopida, hind wings. A) A. suturalis, macropterous wing; B) A. impressa brachypterous wing.

FIGURE 7. Atopida, male terminal segments and genitalia (A, C) and gut contents (B). A) A. suturalis, B) same, gut contents, C) A. paparoa sp. nov.

FIGURE 8. Atopida, abdominal segments of females. A) A. castanea; B) A. hirta; C) A. impressa; D) same, tergite VII; E) A. insularis sp. nov.; F) A. montana, male; G) A. paparoa sp. nov.; H) A. proba; I) same, tergite VII; J) A. suffusa; K) A. suturalis; L) same, tergite VII; M) A. tuhua; N) A. villosa; O) A. westlandensis sp. nov. Not to the same scale.

FIGURE 10. Atopida, bursal sclerites, posterior portion on the right side of the photographs. A) A. castanea; B) A. hirta; C) A. impressa; D) A. insularis sp. nov.; E) A. montana; F) A. paparoa sp. nov.; G) A. proba; H) A. suffusa; I) same; J) A. suturalis, lateral view; K) same, dorsal view; L) A. tuhua sp. nov.; M) A. villosa; N) same; O) A. westlandensis sp. nov. Not to the same scale.

FIGURE 15. Atopida, male genitalia, penis (left), tegmen (right). A) A. proba, B) A. suffusa, C) A. suturalis, D) A. tuhua sp. nov. Scale bars = 0.1 mm.

FIGURE 17. Atopida, male genitalia: penis (left) and tegmen (right), arranged according to overall similarity. A) A. castanea, B) A. suturalis, C) A. proba, D) A. hirta, E) A. montana, F) A. villosa, G) A. westlandensis sp. nov., H) A. impressa, I) A. suffusa, J) A. waipoua sp. nov., K) A. walkerae sp. nov., L) A. lescheni sp. nov., M) A. paparoa sp. nov., N) A. paringa sp. nov., O) A. insularis sp. nov., P) A. tuhua sp. nov. Not to the same scale.

FIGURE 18. Atopida, distribution. A) A. castanea, B) A. hirta, C) A. impressa, D) A. insularis sp. nov., E) A. lescheni sp. nov., F) A. montana, G) A. paparoa sp. nov., H) A. proba, I) A. suffusa, J) A. suturalis, K) A. villosa, L) A. paringa sp. nov., A. tuhua sp. nov., A. waipoua sp. nov., A. walkerae sp. nov. and A. westlandensis sp. nov.