Balticeler Schmidt & Maddison, 2021
|
publication ID |
https://dx.doi.org/10.3897/dez.68.66181 |
|
publication LSID |
lsid:zoobank.org:pub:0D454AFB-9C31-42C3-A5B9-CD9E256C1F41 |
|
persistent identifier |
https://treatment.plazi.org/id/A33BE760-1C69-4DE3-BEE5-E2168C3CCCA2 |
|
taxon LSID |
lsid:zoobank.org:act:A33BE760-1C69-4DE3-BEE5-E2168C3CCCA2 |
|
treatment provided by |
|
|
scientific name |
Balticeler Schmidt & Maddison |
| status |
gen. nov. |
Balticeler Schmidt & Maddison gen. nov.
Type species.
Balticeler kerneggeri sp. nov.
Description.
Diagnosis: Small, markedly shiny ground beetle (due to reduced microsculpture on body surface), with nearly cylindrical body shape (Figs 11-14 View Figures 11–14 ); pronotal marginal borders absent, very slender mandibles and maxillae, and with basal protarsomeres in male not widened.
Head: Slender in its anterior part, robust from level of eyes towards base, with disc markedly convex, base broad and neck constriction absent (Figs 1 View Figures 1–7 , 15 View Figures 15–20 ). Mandibles and maxillae notably long and slender, teeth on mandibular internal margin small, not prominent, markedly shifted basally, mandibular scrobe with seta (Fig. 17 View Figures 15–20 ). Labrum with apical margin very slightly concave, with six setae near apical margin (Fig. 1 View Figures 1–7 ). Clypeus with two primary setae on each side (Fig. 15 View Figures 15–20 ). Anterior and posterior tactile supraorbital setae present (Figs 1 View Figures 1–7 , 15 View Figures 15–20 , 16 View Figures 15–20 ); suborbital seta absent. Furrows on head disc shallow, very short, terminating posteriorly at level of anterior supraorbital setae (Figs 1 View Figures 1–7 , 15 View Figures 15–20 ). Eyes moderately large, moderately protruded laterally; tempora short, about quarter of eye length, very slightly wrinkled towards the neck (Figs 15 View Figures 15–20 , 16 View Figures 15–20 ). Antennae moderately slender, pubescent beginning from second antennomere; pedicellus about as long as scapus and third antennomere (Fig. 3 View Figures 1–7 ). Ligula long and slender, with apex pointed, partly fused with paraglossae, latter distinctly protruded apically (Fig. 19 View Figures 15–20 ); chaetotaxy of ligula not visible with the methods we used. Mentum with median tooth simple, with one pair of sharply defined pits; mentum and submentum separated by distinct suture; mentum with one pair of tactile setae laterally of apical tooth, submentum with two pairs of tactile setae in normal position (Fig. 18 View Figures 15–20 ). Terminal maxillary and labial palpomeres about as long as penultimate palpomere, with shape slightly conical (Fig. 1 View Figures 1–7 ); penultimate maxillary palpomere glabrous; penultimate labial palpomere glabrous except for two long setae in middle.
Prothorax: Small, subcordate, with disc markedly convex, and with apical, basal and lateral borders absent (Figs 13 View Figures 11–14 , 14 View Figures 11–14 , 20 View Figures 15–20 ; in CT images from some angles, such as that shown in Fig. 20 View Figures 15–20 , the shrunken exoskeleton and the negative imprint of the beetle on the inclusion wall may give the impression of existing border lines); anterior margin straight with lateral angles rounded, not protruded anteriorly; basal margin slightly convex in middle and with outer sixth markedly shifted anterad; laterobasal angles moderately large, obtuse, very slightly protruded laterally; anterior lateral seta situated near anterior end of second pronotal third, posterior seta situated at basal angle (Figs 8 View Figures 8–10 , 20 View Figures 15–20 ). Median line on pronotal disc deep between anterior and posterior transverse impressions, absent near apex and base; anterior transverse impression distinct, complete; posterior transverse impression broad, marked by three large pits medially, and small, irregularly impressed laterobasal grooves laterally (Fig. 20 View Figures 15–20 ). Prosternum coarsely punctate (Figs 12 View Figures 11–14 , 22 View Figures 21–25 ); prosternal process short, widened towards truncated posterior margin (Fig. 22 View Figures 21–25 ); procoxal cavities closed externally.
Pterothorax: Elytra in dorsal view moderately ovate to sub-parallel, much broader than pronotum, with sides very slightly narrowed towards broad humerus (Figs 3 View Figures 1–7 , 8 View Figures 8–10 , 11 View Figures 11–14 , 26 View Figures 26–28 , 29 View Figures 29–31 , 36 View Figures 36–42 , 38 View Figures 36–42 , 41 View Figures 36–42 ), in lateral and frontal or caudal views markedly convex towards disc, such that hind body is almost circular in cross-section (Figs 9 View Figures 8–10 , 13 View Figures 11–14 , 14 View Figures 11–14 , 21 View Figures 21–25 , 28 View Figures 26–28 , 30 View Figures 29–31 , 37 View Figures 36–42 , 39 View Figures 36–42 , 40 View Figures 36–42 , 42 View Figures 36–42 ). Basal border absent from base of 5th stria inwards (Fig. 11 View Figures 11–14 ). Parascutellar stria abbreviated, connected with the first stria, parascutellar tactile seta present, situated at base of second stria (Fig. 11 View Figures 11–14 ). Striae 1-9 slightly impressed in anterior 4/5 but markedly accented by rows of large and deeply engraved punctures (Figs 3 View Figures 1–7 , 8 View Figures 8–10 - 11 View Figures 11–14 , 21 View Figures 21–25 , 29 View Figures 29–31 , 36 View Figures 36–42 , 41 View Figures 36–42 ); punctures gradually less deep from anterior 1/9 towards base and posterior quarter towards apex, latter smooth beside the deeply engraved 1st and 8th striae, with recurrent stria absent and subapical setiferous pore “isolated” (the seventh and the internally adjacent striae are invisible) (Figs 2 View Figures 1–7 , 21 View Figures 21–25 ); intervals convex on elytral disc, flat near elytral base and apex. Three discal setae present on each elytron; all are connected to the third stria, with anterior one located near end of anterior elytral 5th, second located slightly anterior of elytral middle, and posterior one located slightly posterior of elytral 2/3 (Figs 9 View Figures 8–10 , 11 View Figures 11–14 ). Umbilicate series consists of eight setae distinctly separated into three groups (Fig. 14 View Figures 11–14 ): humeral group (four setae, with posterior one slightly more distant than setae 1-3), medial group (two setae, located distinctly behind elytral middle), subapical group (two setae). Elytral subapical plica present (Fig. 21 View Figures 21–25 ). Hindwings fully developed. Mesoventrite and metaventrite near anterior margin coarsely punctate (Figs 12 View Figures 11–14 , 22 View Figures 21–25 ); mesocoxal cavities conjunct; mesepimeron wide, metanepisternum long and slender, external lobe of metepimeron well developed (Fig. 22 View Figures 21–25 ).
Legs moderately robust, short (Fig. 12 View Figures 11–14 ). Protibia with antenna cleaner anisochaetous Grade B ( Hlavac 1971), with basal portion of cleaning channel flat, extending basad far beyond insertion of the clip setae, and with insertion of posterior spur very slightly distad of that of the clip setae (Figs 24 View Figures 21–25 , 25 View Figures 21–25 ); protibial external surface smooth, without longitudinal groove, and with apicolateral surface obliquely excised (Fig. 23 View Figures 21–25 ). Basal protarsomeres of males not dilated, not uniquely dentate (Fig. 23 View Figures 21–25 ). Fifth tarsomeres of all legs ventrally smooth, without setae. Mesocoxa with one seta at external margin; metacoxae laterally not extended to elytral epipleuron, trisetose, with both the external setae distinctly removed from the coxal ridge; metatrochanter with a single seta (Fig. 22 View Figures 21–25 ).
Abdomen: Smooth beside primary setation: segments IV-VI with a single, VII with two pairs of setae near apical margin (Fig. 2 View Figures 1–7 ).
Male genitalia: Shape of the parameres of Trechitae type, nearly symmetrical, in general structure similar to parameres of Patrobini , markedly large, each with a long and slender apical apophysis which is more strongly sclerotized on its internal margin, with a large and almost discoidal middle portion, and with a heavily sclerotized basal portion (Figs 32a-d View Figures 32–35 , 35a View Figures 32–35 ); chaetotaxy and membranous parts (e.g., those which probably connect apical and medial portions of the parameres externally) not recognizable. Median lobe moderately long and slender, in lateral view slightly sinusoidal in apical third (Figs 32a View Figures 32–35 , 33 View Figures 32–35 , 34a View Figures 32–35 ), in dorsal view with its distal portion very slightly bent to the right (Figs 32c, d View Figures 32–35 , 34b View Figures 32–35 ); apical lamella well-developed, almost as broad as median lobe. Dorsal surface of median lobe completely open: basal and apical ostia broadly connected across the dorsal median lobe surface, with separated lateral lobes of median lobe basal bulb (Figs 32b, c View Figures 32–35 ).
Etymology.
The generic name compounds the geographical term “Balticum” which is the origin of the amber where the new fossil lineage is preserved, and the Latin verb “celare” (concealing), and therewith refers to the specific circumstance that an odd lineage of ground beetles is hidden in Baltic amber.
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.