Tomarus roigjunenti Neita and Ratcliffe, 2017

Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 8-10

publication ID

https://doi.org/ 10.5281/zenodo.5172580

publication LSID

lsid:zoobank.org:pub:F8317A32-DFD7-42EA-8BC6-DCB5A3659F6C

persistent identifier

https://treatment.plazi.org/id/A24087A7-396A-0427-FF7F-FB71FA32FF4C

treatment provided by

Felipe

scientific name

Tomarus roigjunenti Neita and Ratcliffe
status

sp. nov.

Tomarus roigjunenti Neita and Ratcliffe , New Species

( Fig. 1e–f View Figure 1 , 4e View Figure 4 , 5e View Figure 5 , 6i–j View Figure 6 , 7e View Figure 7 , 8e View Figure 8 , 10i–j View Figure 10 , 11e View Figure 11 , 12e View Figure 12 , 13e View Figure 13 , 14e View Figure 14 , 15e View Figure 15 , 17a View Figure 17 )

Holotype male labeled: “VATRA LAUFQUEN / 14580 M.S.N.M/Neuquén-Arg. / 13-XI-69 / Leg. M. Gentili ”, and with our red holotype label [ IADIZA] . Allotype female labeled: “ R. A. Neuquén / 5 km. S de Lag. Blanca / 15/01/2007 / G. San Blas Col. / 39°06´56.1´´S/70°00´10.1´´W. / 1975 msnm”, and with our red allotype label [ IADIZA] . Paratypes labeled: “Buenos Aires: Prov. de / Buenos Aires / J. Boso Col.” (3 males, 1 female) [ MLPA]. “Buenos Aires” (5 males, 5 females) [ MLPA]. “Chubut: Chubut” (1 male) [ MACN]. “Mendoza: RA. Mendoza / Uspallata / Enero 1975 / S. Roig” (1 male) [ IADIZA]. “Mendoza / Tunuyan / I-1940 ” (1 male) [ MLPA]. “Neuquén: Neuquén / Covunco / 19-XII-1963 ” (1 male) [ IADIZA]. “R. A. Neuquén / Barrancas / 5-1970 Col. A. Roig / CE 00283” (1 female) [ IADIZA]. “Covunco / Neuquén-Arg. / 29-X-62 / Leg. M. Gentili” (1 male) [ IADIZA]. “Neuquén / L. Victoria / Febr. 1958” (1 female) [ IADIZA]. “Neuquén / Covunco / 19-XII-1963 ” (1 male) [ IADIZA]. “ 8-XII-1955 / Collón Cura/ Pcio. Neuquén / Leg. S. Sochajovskoy” (2 females) [ MLPA]. “Río Negro: Arg. Río Negro /Gral. Rocas / Cnia. Catriel 311 m / S. Roig / G. Flores /04/I/05 // 37°52´83´´ S/67°50´16´´W” (3 males, 1 female) [IA- DIZA]. “REP. ARGENTINA / Río Negro / H. Ritcher” (7 males, 1 female) [ MLPA]. “ III-5-1929 / S.O. / Río Negro” (1 female) [ MACN]. “Río Negro / Allen” (1 female) [ MACN]. No Data (3 females) [ MACN]. “Col. Breyer” (1 male) [ MACN]. “REP. ARGENTINA / A. BREYER / Col. Breyer” (1 male) [ MACN]. “ Neuquen / Capital / i.iii.961 // Argentina / 1968 colln. / J. Daguerre” (1 male, 1 female) [ USNM]. “21- II-60 / Cipolletti / Prov Río Negro / lg. S. Schajovskoy / Howden coll. ex. / A. Martínez coll.” (4 males, 2 females) [ CMNC, BCRC]. “ 24-XI-1955 / Villa Regina / Pcia. Rio-Negro / Lg. E. Fleiss // Howden coll. ex. / A. Martínez coll. // Lgyrus [sic] (T.) / gyas / niger / (Blanch.) / A. MARTÍNEZ-DET. 1990” (1 male) [ CMNC]. Paratypes each with our yellow paratype label.

Description of Holotype. Length 26.8 mm; width 13.0 mm. Color dark reddish brown to black. Head: Frons and clypeus coarsely rugose. Frontoclypeal region with 2 distinctly transverse tubercles; tubercles broadly separated, usually about 3 tubercle diameters apart ( Fig. 4e View Figure 4 ). Clypeus strongly rugopunctate to transversely rugose, tapering to slightly emarginate, broad apex with 2 reflexed teeth. Mandibles with 2 teeth, apical tooth subacute, subapical notch narrow ( Fig. 6i–j View Figure 6 ); labium with lateral edge densely setose, paraglossa developed, apex slightly concave at center ( Fig. 7e View Figure 7 ); maxilla with galea developed and with 6 teeth ( Fig. 5e View Figure 5 ). Pronotum: Surface on disc moderately punctate, sides and anterior angles densely punctate to rugopunctate; punctures moderate in size on disc, becoming larger laterally; fovea rugose within. Anterior margin with a strong, median tubercle; fovea small, narrower than distance between eyes ( Fig. 8e View Figure 8 ). Elytra: Surface densely punctate; punctures mostly moderately in size with sparser, minute punctures between larger punctures; larger punctures usually ocellate, some in double rows and with 3 pairs of double rows. Pygidium: Surface densely punctate, punctures becoming denser in basal and lateral angles; punctures small to moderate in size, weakly ocellate; in lateral view with convex surface. Legs: Protibia tridentate and with weak, basal convexity suggestive of 4 th tooth, teeth subequally spaced from each other. Protarsus simple, not enlarged. Mesotibia on transverse carinae with spinules densely (subcontiguous) to broadly (separated by 1 spinule diameter) spaced ( Fig. 10i–j View Figure 10 ). Metatibia with 1 basal, transverse carina interrupted at middle (carina with spine-like setae) and 1 medial, transverse carina at middle; apex with about 21 spinules ( Fig. 10j View Figure 10 ). Venter: Prosternal process long, thick, apex longitudinally oval. Mesosternal surface glabrous, slightly concave at middle. Metasternal surface with sparse, ocellate punctures. Male Genitalia: Parameres ( Fig. 11e View Figure 11 , 12e View Figure 12 , 13e View Figure 13 ) subtriangular, shaft strongly narrowed to apices, mid-shaft with large, curved tooth extending laterally. Internal sac with 3 copulatory lamellae ( Fig. 15e View Figure 15 ).

Allotype Female. Length 27.5 mm; width 13.9 mm. Similar to male, but pronotal surface with punctures slightly less dense, pygidial surface weakly convex. Gonocoxite larger than gonocoxal sternite, with a semicircular furrow (14e).

Paratypes. Length 26.3–27.5 mm; width 12.9–13.9 mm. The 32 male and 20 female paratypes do not significantly differ from their respective primary type other than slight differences in size.

Etymology. This species is named in honor of Dr. Sergio Roig-Juñent in recognition of his invaluable entomological research in Argentina.

Distribution. Tomarus roigjunenti occurs in the provinces of Mendoza, Neuquén, and Río Negro in Argentina.

Locality Records ( Fig. 17a View Figure 17 ). 52 specimens examined (see type information above).

Temporal Distribution. January (7), February (5), March (4), May (1), October (1), November (2), December (3).

Diagnosis. Tomarus roigjunenti is externally similar to T. gyas and T. rosettae . In T. roigjunenti , the apex of the labium is slightly concave ( Fig. 7e View Figure 7 ), while in T. gyas the labium has a small notch ( Fig. 7c View Figure 7 ) and in T. rosettae there is a small tooth ( Fig. 7f View Figure 7 ). The clypeal teeth are distinctly separated in T. rosettae and T. gyas ( Fig. 4c, f View Figure 4 ) but nearly contiguous in T. roigjunenti ( Fig. 4e View Figure 4 ). The tubercles on the head are relatively close together in T. maimon , T. rosettae , and T. roigjunenti ( Fig. 4d–f View Figure 4 ) but widely separated from each other in T. gyas ( Fig. 4c View Figure 4 ). The pronotal fovea is narrow or nearly obsolete in T. maimon , T. rosettae , and T. roigjunenti ( Fig. 8d–f View Figure 8 ) but nearly round and usally as wide as the distance between the eyes (rarely less) in T. gyas ( Fig. 8c View Figure 8 ). The protibia has three teeth in T. maimon ( Fig. 9b View Figure 9 ) and 4 teeth in T. gyas , T. rosettae , and T. roigjunenti ( Fig. 9a View Figure 9 ). The parameres are different among the species (compare Fig. 11c View Figure 11 , 12c View Figure 12 , 13c View Figure 13 ; 11d View Figure 11 , 12d View Figure 12 , 13d View Figure 13 ; 11e View Figure 11 , 12e View Figure 12 , 13e View Figure 13 , and 11f View Figure 11 , 12f View Figure 12 , 13f View Figure 13 ). The internal sac of T. maimon , T. roigjunenti and T. rosettae has three copulatory lamellae ( Fig. 15d–f View Figure 15 ), but in T. gyas there are five lamellae ( Fig. 15c View Figure 15 ).

Life History. Adults are attracted to lights at night. Some specimens have been taken in sand dunes.

MACN

Museo Argentino de Ciencias Naturales Bernardino Rivadavia

USNM

Smithsonian Institution, National Museum of Natural History

BCRC

Bioresource Collection and Research Center

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Dynastidae

Genus

Tomarus

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