Helonoton foleyi Lord and Ivie, 2016

Lord, Nathan P. & Ivie, Michael A., 2016, Several New Genera and Species of New World Synchitini (Coleoptera: Zopheridae: Colydiinae), The Coleopterists Bulletin 70 (4), pp. 715-753 : 715-753

Helonoton foleyi Lord and Ivie , new species ( Figs. 11 , 24 )

Diagnosis. This species is most similar to H. tico in general appearance and possessing narrow, elongate carinae at the base and apex of the elytra, but can be distinguished by the presence of a tubercle/short carina on interstrial interval 7 at the elytral mid-point, the central carinae on pronotal disc less strongly raised, similar to the anterior and posterior carinae on the disc, and the Panamanian distribution.

Description. Size moderate (TL = 3.4 mm), body elongate (TL/EW = 2.2), parallel-sided; light brown to dark brown; habitus as in Fig. 11 . Head: Elongate (HW = 0.6 mm), not constricted behind eyes. Antennae: 11-segmented, ending in an abrupt, 2-segmented club; antennomeres 10–11 forming distinct club, antennomere 10 transverse, trapezoidal to slightly asymmetrical, narrowest at base; antennomere 11 truncate at base, transverse, rounded apically. Prothorax: Pronotum subquadrate to elongate (PW = 1.1 mm, PL = 0.9 mm, PL/PW = 0.8), widest at apical ¼, narrowest at base. Lateral margins of pronotum sinuate, widening apically, anterior ⅓ produced into small arcuate lobe, lateral margin with constriction at middle, posterior ½ subtly sinuate, microdenticulate; anterior angles distinct, angulate, slightly projecting forward to about level of anterior margin; posterior angles not apparent, evenly rounded. Pronotal disc with complex patterns of ridges and depressed areas; central portion with depressed area, bordered laterally by sinuate ridge, ridge equally raised throughout; central portion bordered anteriorly by paired, short, parallel ridges that become confluent with anterior margin; basal portion of central discal area with paired, basolaterally directed ridges that end before posterior margin creating subtriangular baso-medial depression. Pair of short, mid-lateral carinae present between lateral pronotal margin and basal 1/2 of central sinuate tuberculate ridge, as well as smaller pair of shorter carinae present between lateral pronotal margin and anterior portion of central sinuate tuberculate ridge. Procoxal cavities broadly open externally. Mesothorax: Scutellum well-developed, visible, ovoid to pentagonal; anterior margin rounded to truncate, posterior margin rounded to slightly angulate. Elytra: Elongate, parallel-sided, widest at middle (EL = 2.5 mm, EW = 1.5 mm, EL/EW = 1.7). Surface with elongate tubercles or short carinae. Sutural stria raised, beaded, abruptly diverging antero-laterally to form scutellary striole. Each elytral base with 3 elongate carinae; 1 st at base of elytral interstrial interval 3, raised, elongate, distinctly swollen posteriorly; 2 nd at base of interval 5, ½ as long as carina on interval 3; 3 rd at base of interval 7, beginning at humeral angle and extending towards apex, 2X as long as carina on interval 5. Interval 7 in middle of elytra with distinct tubercle. Interval 9 without distinctly raised carina. Remaining tubercles on middle portion of elytra small and round to elongate-oval, subequal in size. Carinae present on elytral declivity, long, extending nearly to apex. Epipleuron present, weakly defined, incomplete to apex, ending at abdominal ventrite IV. Metaventrite: Longer than abdominal ventrite I, with paired sinuate grooves posteriad and bordering mesocoxae, also with paired, slightly curved grooves anteriad and bordering metacoxae. Discrimen moderate, extending to middle ½ of metaventrite, strongly impressed between and anteriad metacoxae. Metacoxae transverse, narrowly separated. Metendosternite of fully winged form as in Fig. 2g (see genus-group description). Abdomen: Intercoxal process of abdominal ventrite I triangular, apex acute. In males, ventrites III and IV with large, smooth, transverse-oval patches in lateral areas. Metathoracic wing: Fully developed.

Distribution. Panama ( Fig. 24 ).

Biology. Specimens were taken on funguscovered logs.

Etymology. Named after Ian Foley (Helena, MT, USA) for his camaraderie and numerous contributions to this paper.

Type Material. 4 specimens. Holotype, male (pointed, partially dissected, genitalia and abdomen in glycerine in genitalia vial pinned underneath specimen, CNCI): PAN: Chiri ; 1700m; 2km W Cerro Punta; Audubon For. 19.v.77; S&J.Peck, Ber 366 . Paratype, female (pointed, partially dissected, genitalia and abdomen in glycerine in genitalia vial pinned underneath specimen, head and prothorax glued to point, CNCI): PANAMA, Prov. Chiriqui; Cerro Punta , 8000 ft.; July 18-24, 1961; J. M. Campbell . Paratype (pointed, KSEM): PANAMA: Chiriqui Prov.; La Fortuna ,”Río Hornito; Trail.” 08°42′N, 82°14′W; 1000 m, 11 VI 1995; J. Ashe & R. Brooks #175; ex: logging fungusy log . Paratype (pointed, KSEM): PANAMA: Chiriquí; 27.7 km W Volcan,; Hartmann’ s Finca , 1650-1700 m; 8°51′48″N, 82°44′36″W; 18 Jun 1996; J. Ashe,; R.Brooks PAN1AB96 194; ex: fungusy log / [label with barcode] SM0050612; KUNHM-ENT .

Figures

Figs. 9–12. Helonoton species, dorsal habitus. 9) H. chiriqui, holotype (KSEM); 10) H. costaricense, holotype

Figs. 23–24. Distribution maps of Helonoton species. 23) Species in Mexico, Guatemala, El Salvador, and Honduras;

Fig. 2. Helonoton costaricense. a) Head, ventral view, b) Labium, ventral view, c) Maxilla, ventral view, d) Antenna, e) Fore leg, f) Mandible, ventral view, g) Metendosternite, ventral view, h) Prothorax, ventral view, i) Pterothorax, ventral view, j) Abdominal ventrites I–V, ventral view, k) Aedeagus, dorsal view, l) Aedeagus, lateral view.