Scatella (Scatella) septemfenestrata Lamb

Zatwarnicki, Tadeusz & Irwin, Anthony G., 2018, Taxonomic notes on the genera Scatella and Scatophila (Diptera: Ephydridae) with a remark on Trixoscelis chilensis (Trixoscelididae), Zootaxa 4377 (1), pp. 91-109: 98-100

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Scatella (Scatella) septemfenestrata Lamb


Scatella (Scatella) septemfenestrata Lamb 

( Fig. 10–11View FIGURES 10–11)

Scatella septemfenestrata Lamb, 1912: 328  .

Scatella septempunctata Malloch, 1933a: 8  , new synonym. Scatella gratiellae Canzoneri & Raffone, 1987: 58  , new synonym.

Type material examined or photographed: The female lectotype of the senior synonym ( Fig. 10View FIGURES 10–11), designated by Mathis and Zatwarnicki (2003), is labelled: "Type H.T. [round label with a red border]/Seychelles Is. Prof. J. S. Gardiner. 1914-537/TYPE [blue label glued to the larger determination label] Scatella septemfenestrata  . Lamb det CGL [except for the "TYPE" label, handwritten]." The lectotype is glued to a rectangular card by its left side (card also has the number "108" handwritten on top; on the underside of the card, a label with "Mahe, '08-9. Seychelles Exp." is glued), is in fair condition (left side obscured by card and glue), and is in the NHMUK. In addition, there is a second female, its head is missing, with the same locality label. The holotype of S. septempunctata Malloch  is labelled: “Vaituha Eiao”, “ X-3-29 200 ft ”, “Marquesas Islands”, “A. M. Adamson collector”, “Pacific Entomological Survey”, “ Scatella septempunctata  Type det JRMALLOCH 1180” (with black submarginal line; det JRMALLOCH printed), and 22 paratypes: ten specimens labelled: “Vaituha Eiao”, “ X-3-29 200 ft ”, “Marquesas Islands”, “A. M. Adamson collector”, “Pacific Entomo- logical Survey”, “Paratype” (yellow) ( Fig. 11View FIGURES 10–11); five specimens labelled: “Hanamenino Valley Sea level, VII.17.1930”, “Tahuata Marquesas Is.”, “LeBronnec & H. Tauraa, Coll.”, “Pacific Entomo- logical Survey”; six specimens labelled: “Oomoa [= Omoa] Valley Near sea-level VIII.21.1930”, “Fatuhiva Marquesas Is.”, “LeBronnec Collector”, “Pacific Entomo- logical Survey”, and one specimen labelled: “Tahauku VII - 10-1929 ”, “Marquesas Is. Hivaoa”, “Mumford & Adamson”, “Pacific Entomological Survey” preserved in BPBM. The holotype of S. gratiellae  is labelled: "KENYA L. NAIVASHA 14. VII. 79 W. ROSSI" (handwritten), "HOLOTYPUS ♀ scatella gratiellae  nov. det. Canzoneri S. & G. Raffone" (red, name and ♀ handwritten), and one paratype bearing the same locality label, preserved in MSNVE.

Material examined: AUSTRALIA: Norfolk Island, shade house in plant nursery, 2. XII. 2014, R  . A. Broadley, 1 ♂ (AGI)  ; Queensland, Hermit Park, malaise trap, 1. VII. 2015, G. Cocks, 1 ♂ (AGI). KENYA  : Tsavo National Park (West), 4 –7. IV. 1968, B. Cogan & A. Hutson, 1 ♂ ( NHMUKAbout NHMUK). MALAYSIA  : Malay Peninsula, Pahang, F.M.S. Gunong Jahan Padang 5550', H. M. Pendlebury, 11. XII. 1922, 1 ♀  ; 5. I. 1923, 1 ♂ ( NHMUKAbout NHMUK). MAURITIUS  : La Re'duit, 26. V. 2000, J. W. Ismay, 3 ♂ and 50 in plastic container; Plaine Lievre   , 28. V. 2000, J. W. Ismay, 2 ♀ (NMWC); Rodrigues I.  , VIII–XI.1918, H. J. Snell & H. P. Thomasset, 1 ♂ ( NHMUKAbout NHMUK). NAMIBIA  : Ameib Farm (W. 30), on vegetation around pools, 31. I.–2. II. 1972, South Western Africa Expedition, 1 ♂ ( NHMUKAbout NHMUK)  ; Namib Desert Park, Kuiseb River at Gobabeb, 400 m, Riverine forest and sand, 12. II. 1974, ME & BJ Irwin, 2 ♂, 6 ♀ ( NMSAAbout NMSA)  ; L. Lyneborg, 1 ♂ ( NHMUKAbout NHMUK). NIGERIA  : Lake Chad . Mallam Fatori , 13. IV. 1967, J. C. Deeming, 1 ♀ ( NHMUKAbout NHMUK). REPUBLIC OF SOUTH AFRICA: Kwa Zulu Natal: Kosi Bay Nature Res. (82 m), Malaise trap, woodland, savanna, 26°57.17'S, 32°49.27'EGoogleMaps  , 13.–17. XII. 2010, A  . H. Kirk-Spriggs, 1 ♀; Ndumo Game Reserve, Malaise trap, grassy foodplain, 26°54.288"S, 32°17.974"E, 9.–10. XII. 2009, AGoogleMaps  . H. Kirk-Spriggs, 1 ♀; Ndumo Game Reserve, Skokwe area, Ficus forest , 26°52.125'S, 32°13.731'E, 37 m, 30. XI.–4. XII. 2009, AGoogleMaps  . H. Kirk-Spriggs, 1 ♂, 1 ♀; Umlazai Nat. R., 10 m, sweeping, dune forest, 28°57'40"S, 31°46'04"E, 17. XII. 2010, AGoogleMaps  . H. Kirk-Spriggs, 1 ♀; Orange Free State  : Harrismith Dist. , Scotland Farm, Malaise trap, dense Leucosedeadominated scrub, 27°58'59.5"S, 29°37'09.8"E, 10–12. III. 2 0 12, A.H. Kirk-Spriggs, 1 ♀ ( BMSAAbout BMSA)GoogleMaps  . SAINT HELENA. Sud: Sandy Bay, sur pierres pres d'un ruisseau, 1. III. 1967, J. Decelle, N. & J. Lelup, 96 ex; Est: Basse Fisher's Valley , irrigations, 1000 ft  , 14–19. XII. 1965, P. Basilewsky, P.L.G. Benoit, N. Lelup, 176 ex; Est: Basse Fisher's Valley , Bradley Ruins, 1000 ft  , 25. XI. 1965, P. Basilewsky, P.L.G. Benoit, N. Lelup, 12 ex; Basse Fisher's Valley   , 26. I. 1967, J. Decelle, N. & J. Lelup, 3 ex; ( RMCAAbout RMCA); Basse Fisher's Valley   , 6. V. 1967, J. Decelle, N. & J. Lelup, 24 ex ( RMCAAbout RMCA, NHMUKAbout NHMUK). UNITED ARAB EMIRATES  : Al-Ain, Mubazzarah, hot spring, 16. III. 2010, T. Zatwarnicki, 1 ♀; Kalba , beach, 3  . III. 2 0 10, T. Zatwarnicki, 1 ♂ (TZPC); NARC, near Sweihan, light trap, 14–28. III. 2005, A  . van Harten, 1 ♂ ( NMWCAbout NMWC); Wadi Asimah , 435 m  , 8. III. 2010, T. Zatwarnicki, 7 ♂, 11 ♀ ( TZPC); Wadi Maidaq , 390 m  , 3. II. 2010, T. Zatwarnicki, 2 ♂, 1 ♀ ( TZPC); Wadi Shawkah , reservoir 300 m  , 27. II. 2010, T. Zatwarnicki, 3 ♂, 15 ♀ (TZPC).

Type localities: Lamb (1912) reported S. septemfenestrata  from Mahé ( Seychelles): from a marsh just behind the beach at Port Glaud (coordinates: 4°39′S 55°24′E). Type locality of S. septempunctata  is Vaituha, which is a valley on island Eiao (coordinates: 8°S 140.7°W; Marquesas, French Polynesia). Type locality of S. gratiellae  is Lake Naivasha in Kenya (coordinates: 0°46′6.70″S 36°21′2.32″E).

Diagnosis. This species is closely related to S. ciliata Collin  but is distinguished from other species by the wing having six to seven hyaline spots within a dark coloured background (sometimes the apical spot in cell R3 is indistinct), and legs greyish to mostly brownish black; tarsi mostly black.

Distribution. Afrotropical: Ascension Island, Ghana, Kenya, Mauritius, Namibia, Nigeria, Rodriguez, Saint Helena, Seychelles, Sudan, Republic of South Africa (Free State; Kwa Zulu Natal). Australasian/Oceanian: Australia (Queensland), French Polynesia (Marquesas), Norfolk Island. Oriental: Japan (Ryukyu Islands), Malaysia. Palaearctic: United Arab Emirates.

Remarks. Canzoneri & Raffone (1987) compared their new species with the Palaearctic S. stagnalis (Fallén)  and S. subguttata (Meigen)  , but not with S. septemfenestrata  , which was recorded in the same paper. Marginal spots of specimens selected as the new species are less distinct (“slightly lightened” in original description), than typical specimens of S. septemfenestrata  .

In his description of Scatella septempunctata, Malloch  compared his species with S. stagnalis  but did not consider S. septemfenestrata  , which had been described from the Seychelles (over 16,000 km away). Thanks to the kindness of Neal Evenhuis, we were able to compare high quality stacked images of one of the paratypes of Scatella septempunctata Malloch  (collected from Vaituha, Eiao, French Polynesia by Adamson on 3.X.1929). This demonstrated that S. septempunctata  shows no discernable differences from, and falls within the range of variation of S. septemfenestrata  from Australia, the Indian Ocean, Africa and the Atlantic Ocean.


Natural History Museum, London


KwaZulu-Natal Museum


National Museum Bloemfontein


Royal Museum for Central Africa


National Museum of Wales














Scatella (Scatella) septemfenestrata Lamb

Zatwarnicki, Tadeusz & Irwin, Anthony G. 2018


Scatella septemfenestrata

Lamb 1912: 328


Scatella septempunctata Malloch, 1933a : 8

Canzoneri 1987: 58Malloch 1933: 8