Cladophorus humeralis Kleine, 1926

Bocak, Ladislav, 2017, Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea, Zootaxa 4320 (3), pp. 505-522 : 512

publication ID	https://doi.org/ 10.11646/zootaxa.4320.3.6
publication LSID	lsid:zoobank.org:pub:Ca50E5A5-B417-4000-B67D-5F6028Da9154
DOI	https://doi.org/10.5281/zenodo.6018036
persistent identifier	https://treatment.plazi.org/id/A312190F-EC12-FF84-509F-3CCDBF9AFAA3
treatment provided by	Plazi
scientific name	Cladophorus humeralis Kleine, 1926
status

Treatment

( Figs. 10, 24 View FIGURES 6 – 32 , 40 View FIGURES 33 – 52 , 61–62 View FIGURES 53 – 78 )

Cladophorus humeralis Kleine, 1926: 130 .

Type material. ‘ Co-Typus’ [syntype, No. 274]. Male, New Guinea, Bomberai Peninsula , Kapaur ( MIZW). Other material examined. Male, female . Indonesia, West Papua prov., DMP-Wasior, 100–200 m (A00434–35, LMBC).

Diagnosis. C. humeralis has a unique elytral colour pattern with brown humeri and a longitudinal patch in the posterior part of each elytron ( Fig. 40 View FIGURES 33 – 52 ). Further, the phallus of this species is robust and gradually acuminate apically ( Figs. 61–62 View FIGURES 53 – 78 ). C. manokwarensis was identified as a sister species of C. humeralis in the molecular phylogeny ( Fig. 4 View FIGURES 4 – 5. 4 ). Although these species are very close each to other, they differ besides the colour pattern in the shape of the antennomere 3 and the relative size of eyes. The shape of phallus differs only slightly in this pair of species ( Figs. 61–62, 65–66 View FIGURES 53 – 78 )

Redescription. Male. Body large, 15.5–15.7 mm long, dark brown to black including pronotum and substantial part of elytra ( Fig. 40 View FIGURES 33 – 52 ), humeri light brown, bright humeral patches separated by dark coloured area along inner margin of elytra; additionally long, light patch in posterior part of each elytron, posterior patch with gradual transition between dark and light coloured areas of elytron ( Fig. 40 View FIGURES 33 – 52 ). Head small, with hemispherically prominent eyes, frontal interocular distance 1.92 times maximum diameter; antennae long, reaching beyond two thirds of elytral length, flabellate, antennomere 3 with lamella 2.55 times longer than stem, lamella attached to apical part of antennomere ( Fig. 10 View FIGURES 6 – 32 ). Pronotum transverse, 1.43 times wider than long at midline, lateral margins apparently elevated, frontal margin widely rounded, frontal angles inconspicuous, rounded, lateral margins slightly convex, posterior angles prominent, all carinae well-developed ( Fig. 24 View FIGURES 6 – 32 ). Elytra flat, slender, subparallel-sided, transverse costae well-developed, regular, elytral areoles slightly transverse ( Fig. 40 View FIGURES 33 – 52 ). Phallus robust, parallel-sided in middle, with gradually acuminate apex ( Figs. 61–62 View FIGURES 53 – 78 ).

Measurements. BL 15.5–15.7 mm, PL 2.10 mm, PW 3.00– 3.05 mm, WH 1.70–1.75 mm, Edist 1.15 mm, Ediam 0.60 mm, LE 13.2 mm, antennomere 3: stem 1.0 mm, lamella 2.55 mm.

Distribution. Western New Guinea, Bomberai Peninsula, Teluk Wondama regency ( Fig. 3 View FIGURES 1 – 3. 1 – 2 )