Enicospilus kakanicus Shimizu, 2020

Shimizu, So, 2020, The Nepalese species of the genus Enicospilus Stephens, 1835 (Hymenoptera, Ichneumonidae, Ophioninae): a preliminary revision and identification key to species, Deutsche Entomologische Zeitschrift 67 (1), pp. 69-126 : 69

publication ID

https://dx.doi.org/10.3897/dez.67.51332

publication LSID

lsid:zoobank.org:pub:2B601B5D-E1BD-44B7-BA89-554E3AB5EAE1

persistent identifier

https://treatment.plazi.org/id/CF0FE094-738D-4491-8EBA-3C673E97C238

taxon LSID

lsid:zoobank.org:act:CF0FE094-738D-4491-8EBA-3C673E97C238

treatment provided by

Deutsche Entomologische Zeitschrift by Pensoft

scientific name

Enicospilus kakanicus Shimizu
status

sp. nov.

Enicospilus kakanicus Shimizu sp. nov. Figs 2F View Figure 2 , 13 View Figure 13

Etymology.

The specific name is derived from the type locality, Kakani, Nepal.

Material examined.

1♂: Nepal .

Type series: holotype ♂, Kakani (2,000 m), Nepal, VIII.1982, M.G. Allen leg. (LT) ( NHMUK) (Figs 2F View Figure 2 , 13 View Figure 13 ) .

Distribution.

Nepal.

Description.

Male (Holotype) (Fig. 13 View Figure 13 ). Body length ca 24.0 mm.

Head with GOI = 2.8 (Fig. 13C View Figure 13 ). Lower face 0.7 × as wide as high, moderately punctate with setae, strongly shiny (Fig. 13B View Figure 13 ). Clypeus 1.5 × as wide as high, moderately punctate with setae, moderately convex in profile, and its lower margin impressed (Fig. 13B, C View Figure 13 ). Malar space 0.4 × as long as basal mandibular width (Fig. 13B, C View Figure 13 ). Mandible moderately twisted by ca 30°, moderately long, more or less evenly tapered, its outer surface with a diagonal setose groove between its dorsoproximal corner to base of mandibular apical teeth (Fig. 13B, C View Figure 13 ). Upper mandibular tooth 2.0 × as long as lower one (Fig. 13B View Figure 13 ). Frons, vertex and gena strongly shiny with fine setae (Fig. 13B-D View Figure 13 ). Posterior ocellus close to eye (Fig. 13B-D View Figure 13 ). Ventral end of occipital carina joining oral carina. Antenna with 73 flagellomeres; first flagellomere 1.7 × as long as second; 20th flagellomere 2.8 × as long as wide.

Mesosoma entirely moderately to strongly shiny with setae (Fig. 13E View Figure 13 ). Pronotum punctate with wrinkles centrally. Mesoscutum 1.4 × as long as its maximum width, finely punctate with setae, and evenly rounded in lateral profile (Fig. 13E View Figure 13 ). Notauli absent (Fig. 13E View Figure 13 ). Scutellum moderately convex, finely punctate with setae, with lateral longitudinal carinae reaching anterior 0.6 (Fig. 2F View Figure 2 ). Epicnemium densely punctate. Epicnemial carina strongly present, evenly and moderately curved to anterior, its dorsal end reaching anterior margin of mesopleuron (Fig. 13E View Figure 13 ). Mesopleuron entirely punctate with longitudinal wrinkles or striae (Fig. 13E View Figure 13 ). Submetapleural carina almost parallel sided (Fig. 13E View Figure 13 ). Metapleuron entirely finely punctate with setae (Fig. 13E View Figure 13 ). Propodeum weakly evenly rounded in profile; anterior transverse carina complete; anterior area longitudinally striate; spiracular area finely punctate with setae; posterior area moderately longitudinally strigose centrally and irregularly wrinkled to reticulate laterally; propodeal spiracle elliptical, its outer margin not joining pleural carina by a ridge (Fig. 13E View Figure 13 ).

Wings. Fore wing length ca 17.0 mm with AI = 0.4, CI = 0.4, DI = 0.3, ICI = 0.5, SDI = 1.4, SI = 0.1, SRI = 0.3; vein 1m-cu&M almost evenly curved; vein 2r&RS straight and RS evenly curved; fenestra and sclerites of discosubmarginal cell as in Figure 13F View Figure 13 ; proximal sclerite triangular, confluent with distal sclerite, strongly pigmented; central sclerite absent; distal sclerite more or less entirely present from base to apex, weakly to moderately pigmented; proximal corner of marginal cell evenly setose; posterodistal corner of second discal cell ca 95°; posterodistal corner of subbasal cell ca 95°; vein 1cu-a antefurcal to M&RS by 0.2 × 1cu-a length (Fig. 13F View Figure 13 ). Hind wing with NI = 1.8, RI = 1.8; vein RS straight; vein RA with six uniform hamuli.

Legs. Outer surface of fore tibia with scattered spines. Hind leg with coxa in profile 1.9 × as long as deep; basitarsus 2.0 × as long as second tarsomere; fourth tarsomere 0.6 × as long as third tarsomere and 4.2 × as long as wide; tarsal claw simply pectinate.

Metasoma with PI = 3.8, DMI = 1.2, THI = 3.6; dorsal margin of tergite 1 not sinuous; thyridium elongate (Fig. 13A View Figure 13 ).

Colour (Fig. 13 View Figure 13 ). Entirely testaceous except for apex of mandible black. Wings hyaline; sclerites of fenestra and veins testaceous.

Variation. Unknown.

Female. Unknown.

Differential diagnosis.

Enicospilus kakanicus sp. nov. is similar to and can be confused with E. longitarsis Tang, 1990, E. tangi sp. nov., and E. yonezawanus (Uchida, 1928). These species all belong to the E. ramidulus complex and share the following characters: outer surface of mandible with a diagonal setose deep groove between its dorsoproximal corner and base of mandibular apical teeth (e.g. Fig. 2B, D View Figure 2 ), fore wing fenestra without central sclerite (e.g. Figs 13F View Figure 13 , 25F View Figure 25 , 27F View Figure 27 ), and proximal sclerite triangular (e.g. Figs 13F View Figure 13 , 25F View Figure 25 , 27F View Figure 27 ). Enicospilus kakanicus sp. nov. is distinguished from the above species by the rather short lateral longitudinal carinae of the scutellum, i.e. reaching the anterior 0.6 of the scutellum in E. kakanicus sp. nov., as in Figure 2F View Figure 2 , but almost always reaching the posterior end of the scutellum in E. longitarsis Tang, 1990, E. tangi sp. nov., and E. yonezawanus , as in, e.g., Figure 2H View Figure 2 , and also by the characters used in the above key, such as width of lower face, mandibular shape and length, and surface sculptures of metapleuron.