Trichocoelina janetscheki ( Lengersdorf, 1953 ) Vilkamaa & Menzel, 2019

Vilkamaa, Pekka & Menzel, Frank, 2019, Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species, Zootaxa 4665 (1), pp. 1-67 : 33

Trichocoelina janetscheki ( Lengersdorf, 1953) comb. n.

Figs 10 A, 10 B , 17 D

Literature. Neosciara janetscheki Lengersdorf—Lengersdorf (1953) : 167, fig. 1; Janetschek (1956): 471. Lycoriella (Hemineurina) janetscheki (Lengersdorf) — Tuomikoski (1959a): 35; Gerbachevskaja-Pavluchenko (1986): 31; Franz (1989): 14; Menzel & Mohrig (2000): 410; Wirta et al. (2016): appendix, unpaginated p. 21 (table S1) and unpaginated p. 39 (cladogram).

Material studied. CANADA, Nunavut, Sverdrup Islands, Elles Ringnes Island, Isachsen , 70.79°N, 103.55°W, lemming burrow, 17.VI.1960, J.R. Vockeroth, 4 males (3 in CNC, 1 in MZH) ; same locality but moss near very small stream, 14.VII.1960, J.F. McAlpine, 2 males (in CNC) ; Nunavut, Ellesmere Island, Fosheim Peninsula, Hot Weather Creek , 79°58’N, 84°28’W, 2.VII.1990, F. Brodo, 1 male (in MZH) ; NE Greenland, Mestersvig , 72.24°N, 23,92°W, C. Vibe (in ZMUC) ; NE GREENLAND, Zackenberg , 74°28’N, 20°34’W (UTM8265500:0513778), 44 m, 2–11.VII.2011, T. Roslin & G. Várkonyi, 1 male (in MZH) ; same locality but UTM 8265758:0513786, 37 m, Malaise trap, 11–20.VII.2011, T. Roslin & G. Várkonyi, 2 males (in MZH, SDEI) ; same locality but 74.50°N, 21.00°W, 44 m, 7.VII.2011, T. Roslin & G. Várkonyi, 1 male (BOLD Sample ID GRPV16, in DAUH) ; same locality but 74.4667°N, 20.5667°W, 48 m, 24.VII.2011, T. Roslin & G. Várkonyi, 3 males (BOLD Sample IDs ZA2012- 50010, ZA2012-50011 and ZA2012-50012, in DAUH) ; USA, Colorado, Mt. Evans , 14 000 ft, 25.VII.1961, W.R.M. Mason, 1 male (in USNM) .

Redescription. Male. Head. Face brown, antenna dark brown maxillary palpus pale yellowish. Eye bridge 2–3 facets wide. Face with 15–21 setae. Clypeus with 1–4 setae. Maxillary palpus with 3 segments, 1 st segment as long as or longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 3–9 setae, with dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.95–2.75x as long as wide, the neck shorter than broad, the longest setae shorter than the width of flagellomere. Thorax. Dark brown, setae pale. Anterior pronotum with 5–11 setae. Proepisternum with 7–24 setae. Scutellum with 4 longer and some short and fine setae.

Wing. Fumose. Length 2.2–3.1 mm. Width/length 0.35–0.40. Anal lobe weak. Veins distinct. c/w 0.50–0.60. R 1 /R 0.50–0.90. stM shorter than fork of M. r-m longer than bM, bM non-setose, r-m non-setose or with 1–2 setae. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a large patch in shallow depression. Fore tibial spur as long as the tibial width. Abdomen. Pale brown, setae pale, short and fine. Hypopygium ( Fig. 10 B ). Brown, as abdomen. Intergonocoxal area long, with two short setose lobes. Gonocoxa broad, as long as gonostylus, medial margin basally smoothly curved, with short and fine setosity. Gonostylus ( Fig. 10 A ) voluminous, apically truncate, impressed; with short setosity, a short apical tooth, and numerous megasetae in the medial impression; megasetae slightly curved or straight, short and slender; with 1 well-differentiated whiplash seta basally at ventromedial margin. Tegmen ( Fig. 17 D ) longer than broad, laterally straight, apically with a narrow hyalinous acuminate process, weakly sclerotized, aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long.

BIN. BOLD:ACK5495.

Discussion. In Fig. 10 A the gonostylus is flattened and distorted to show the arrangement of the megasetae. In its extremely voluminous gonostylus, Trichocoelina janetscheki ( Lengersdorf, 1953) resembles most T. incrassata sp. n., for distinguishing characters, see under the latter.

Figures

FIGURE 10. Trichocoelina janetscheki (Lengersdorf, 1953) (from Greenland). A. Gonostylus, ventromedial. B. Hypopygium, ventral. Scale 0.1 mm.

FIGURE 17. Tegmen, ventral. A. Trichocoelina imitator sp. n. (holotype). B. T. oricillifera sp. n. (paratype from Salla). C. T. hians sp. n. (holotype). D. T. janetscheki (Lengersdorf, 1953) (from Colorado). E. T. nefrens sp. n. (paratype from Rassokha). F. T. magnifica sp. n. (holotype). Scale 0.1 mm.