Asiacercopis Liang, 1994b

Liang, A. - P. & Webb, M. D., 2002, New taxa and revisionary notes in Rhinaulacini spittlebugs from southern Asia (Homoptera: Cercopidae), Journal of Natural History 36 (6), pp. 729-756 : 732-736

Asiacercopis Liang, 1994b . Type species: A. hainanensis Liang

hainanensis Liang, 1994b , S. China

niger (Lallemand, 1927, Au W dus) Laos, Vietnam

nilgiriensis (Lallemand, 1927, Au W dus)? India, Vietnam Circulocercopis gen. nov. Type species: C. ochraceus sp. nov.

fasciatus sp. nov. China, Vietnam, Laos

ochraceus sp. nov. China, Vietnam, Thailand

sinuatus sp. nov. China * Eoscarta Breddin, 1902 ; Liang, 1996a (revision). Type species: E. eos Breddin

Thoodzata Distant, 1908 syn. nov. Type species: T. princeps Distant

Vigilantius Distant, 1916 syn. nov. Type species: V. fascialis Distant

Pseudeoscarta Lallemand, 1933 syn. nov. Type species: P. pendleburyi Lallemand

Eoscartoides Matsumura, 1940 . Synonymized by Liang (1996a). Type species:

E. elongatus Matsumura

Eoscartopsis Matsumura, 1940 . Synonymized by Liang (1996a). Type species:

Monecphora assimilis Uhler

Eoscarta (Paraeoscarta) Lallemand, 1949 . Type species: E. liternoides Breddin

Hemieoscarta Lallemand, 1949 syn. nov. Type species: H. nigripes Lallemand

apicalis (see assimilis )

apicata Distant, 1908 , Sri Lanka

assimilis (Uhler, 1896, Monecphora ) Japan, non-type localities: China, Korea,

Russia

apicalis (Matsumura, 1903) (Rhinaulax) , Japan. Synonymized by

Matsumura, 1904

fusca (Melichar, 1902) ( Cercopis (Callitettix )), Russia. Synonymized by

Kwon and Lee (1979)

atricapilla (see Paracercopis )

aurora Kirkaldy, 1909 , China melli Schmidt, 1920 . Synonymized by Lallemand (1949). China binotata Liang, 1996a , Vietnam borealis (Distant, 1878, Cosmoscarta ) India, non-type localities: China, Burma, Laos, Vietnam tristis (Lallemand, 1927, Cosmoscarta borealis var. tristis ) Laos, syn. nov. elongata ( Matsumura, 1940, Eoscartoides ) Taiwan fascialis (Distant, 1916, Vigilantius ) India, comb. nov. ferruginea Distant, 1916 , India fusca (see assimilis ) fuscata Distant, 1916 , India karschi Schmidt, 1920 , China laoensis (see liternoides ) liternoides Breddin, 1902 , Sumatra, non-type localities: India, Laos, Vietnam, Thailand, Malaysia, Indonesia laoensis Schmidt, 1918. Synonymized by Jacobi (1944). melli (see aurora ) nigripes ( Lallemand, 1949, Hemieoscarta ), W. Malaysia, comb. nov. nilgiriensis (see Asiacercopis ) nobilis Lallemand, 1924 , India ochraceous ( Metcalf and Horton, 1934, Au W dus) China transparena Chou and Wu, 1992. China. Synonymized by Liang (1996a) parva Liang, 1996a , China pendleburyi (Lallemand, 1933, Pseudeoscarta ) Malaya, comb. nov. princeps (Distant, 1908, Thoodzata ) Burma, non-type localities: Malaya, Java, comb. nov. spenceri Liang, 1996a , Cambodia transparena Chou and Wu (see ochraceous ) tonkinensis Lallemand, 1949 , Vietnam, non-type localities: China zonalis ( Matsumura, 1907) (Rhinaulax) , Taiwan

Eoscartoides (see Eoscarta )

Eoscartopsis (see Eoscarta )

Esakius (see Paracercopis )

Horiscarta (see Sounama ( Stenaulophrys ))

Kanozata Matsumura, 1940 . Type species: K. arisana Matsumura arisana Matsumura, 1940 , Taiwan choui ( Yuan and Wu, 1992, Stenaulophrys ) China, comb. nov. contermina (Distant, 1916, Au W dus) India, comb. nov. shillongana (Distant, 1916, Au W dus) India, comb. nov.

Kuscarta (see Sounama ( Stenaulophrys ))

Paracercopis Schmidt, 1925 ; Liang, 1993 (1992) (revision). Type species: Cercopis

(Callitettix) seminigra Melichar

Esakius Ouchi, 1943. Synonymized by Liang (1993) atricapilla (Distant, 1908) (Eoscarta) Burma, non-type localities: China, stat. rev. bicolor Liang, 1993 (1992) China chekiangensis (Ouchi, 1943) (Esakius) China. Synonymized by Liang (1993). stat. rev. transversa Lallemand and Synave, 1955 , China. Synonymized with atricapilla by Liang (1993), syn. nov. crocea (see seminigra var. crocea ) darjilingii (Lallemand, 1924) India, non-type localities: China fuscipennis (Haupt, 1924) China sinensis (Ouchi, 1943) China. Synonymized by Liang (1993) saturata (see seminigra var. saturata ) seminigra (Melichar, 1902) ( Cercopis (Callitettix )) China seminigra var. saturata Haupt, 1924 , China. Synonymized by Liang (1993) seminigra var. crocea ( Metcalf and Horton, 1934) , China. Synonymized by Liang (1993)

Paracercopoides Liang, 1994a . Type species: P. X avithorax Liang assamensis Liang, 1994a , India duplicinigra Liang, 1994a, Vietnam X avithorax Liang, 1994a, China (Hainan Prov.)

Paraeoscarta (see Eoscarta )

Sounama Distant, 1908 . Type species: S. imprimis Distant

Stenaulophrys Jacobi, 1921 . syn nov., stat. nov. Type species: Rhinaulax bimaculatus

Matsumura

Sounama s. str. acuta sp. nov. India imprimis Distant, 1908 , India

Sounama (Stenaulophrys) Jacobi, 1921

Kuscarta Matsumura, 1940 syn. nov. Type species: K. koshunella Matsumura

Horiscarta Matsumura, 1940 . Type species: Rhinaulax bimaculatus Matsumura asiatica sp. nov. Thailand, Burma, Malaya bimaculata ( Matsumura, 1907, Rhinaulax ) Taiwan, comb. nov. borneensis sp. nov. Northern Borneo choui (see Kanozata ) comes (Jacobi, 1921, Thoodzata ) comb. nov. India. coomani sp. nov. Vietnam dentata sp. nov. China expandipenis sp. nov. China horishana ( Matsumura, 1940, Horiscarta ) Taiwan, comb. nov. koshunella ( Matsumura, 1940, Kuscarta ) Taiwan, comb. nov. nigra sp. nov. China robustipenis sp. nov. China, Laos shillongana Distant (see Kanozata )

Stenaulophrys (see Sounama (Stenaulophrys) )

Thoodzata (see Eoscarta )

Vigilantius (see Eoscarta )

Key to the genera of Rhinaulacini from southern Asia

1 Hind wing with Rs and M reduced and jointly united with R near midlength of wing,

i.e. with two closed apical cells (see Liang, 1996a: ®gure 1); subgenital plate with strongly incurved apical process (see Liang, 1996a: ®gure 2)... Eoscarta Breddin ± Hind wing with Rs and M extending independently to apex of wing, united only by a cross-vein near midlength of wing, i.e., with four apical cells (®gure 15); subgenital plate with or without strongly incurved apical process......... 2

2 Hind wing with the cross vein m±cu meeting Cu1a after Cu1a/Cu1b bifurcation and

the basal half of Cu1b strongly arched against Cu2 (see Liang, 1993: ®gure 7); aedeagal shaft robust (®gure 11); subgenital plate with a strongly incurved apical process...................... Paracercopis Schmidt

± Hind wing with m±cu cross vein meeting Cu1a before Cu1a/Cu1b bifurcation and the basal half of Cu1b relatively straight and remote from Cu2; aedeagal shaft elongate (®gure 2) or if robust subgenital plate without a strongly incurved apical process.. 3

3 Hind wing with bifurcation of Cu1a/Cu1b near m±cu cross vein (see Liang, 1994a:

®gures 1, 2); aedeagal shaft robust (see Liang, 1994a: ®gure 7). Paracercopoides Liang

± Hind wing with bifurcation of Cu1a/Cu1b distal of m±cu cross vein (®gure 15); aedeagal shaft elongate (®gure 6)............... 4

4 Fore wings with Rs branched apically (®gure 20); aedeagal shaft very elongate with

®liform coiled or sinuate distal half (®gures 6, 26, 31)... Circulocercopis gen. nov.

± Fore wings with Rs not branched apically (®gures 14, 16, 18); aedeagal shaft not as above..................... 5

5 Aedeagal shaft (®gure 12) with a pair of apical, long, needle-like spines.....

................... Asiacercopis Liang ± Aedeagal shaft without apical spines.............. 6

6 Male pygofer long (®gure 2); subgenital plates lanceolate, caliper-like, basally narrow

with membranous outer margin (®gures 34, 35, 41, 42); small to moderately large species, length 5±8 mm ........... Kanozata Matsumura

± Male pygofer short (®gure 59); subgenital plates not as above; small species, length 4.5±5.8 mm................ Sounama Distant