Spelaeodiscus dejongi Gittenberger, 1969

Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander & Feher, Zoltan, 2018, Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae), ZooKeys 769, pp. 13-48 : 13

Spelaeodiscus dejongi Gittenberger, 1969 Figure 6

Spelaeodiscus (Spelaeodiscus) dejongi Gittenberger, 1969: 295-296, fig. 3.

Spelaeodiscus dejongi - Welter-Schultes 2012: 213.

Type material.

Jama Nadjama bei Gnezdu (Izitovice), Krain, leg. Kuščer, coll. Edlauer, NHMW 49517a (holotype, SW: 2.8 mm, SH: 1.6 mm, Fig. 6A-E ); Same data, NHMW 49517b/17 paratypes. See remarks concerning the type locality.

Other material.

Vetajama bei Sokol, coll. Edlauer, NHMW 48071/2, (det. Gittenberger, 1974); Radetina pećina, Itijino brdo, 1300 m, leg. Dabović, NHMW 48312/11 (large shells, similar to the type); Pećina Marka Vuksanovića, 1400 m, leg. Dabović, coll. Edlauer ex coll. Kuščer 666/10, NHMW 49321/1 (det. Gittenberger, 1974); Montenegro, S of Virpazar, 1 km (in a straight line) ESE of Limljani , near the small road, 323 m a.s.l., 42°11.414'N, 19°06.277'E (site code: 20171019B), leg. T. Deli, Z.P. Erőss, A. Hunyadi & B. Páll-Gergely, 19.10.2017, DT/2, HA/1, HNHM 103198/2, PGB/1 ; Montenegro, S of Virpazar, 0.8 km (in a straight line) NE of Limljani , near the small road, 350 m a.s.l., 42°12.068'N, 19°05.969'E (site code: 20171019C), leg. T. Deli, Z.P. Erőss, A. Hunyadi & B. Páll-Gergely, 19.10.2017, DT/2, HA/2. HNHM 103199/2, PGB/1 ; Montenegro, 1.9 km (in a straight line) S of Virpazar , near the road, 160 m a.s.l., 42°13.312'N, 19°05.440'E (site code: 20171019E), leg. T. Deli, Z.P. Erőss, A. Hunyadi & B. Páll-Gergely, 19.10.2017, DT/24, EZP/24, HA/24, HNHM 103200/24 + 1 photographed shell (Fig. 6K-O ), PGB/24 ; Montenegro, SE of Virpazar, 4.3 km (in a straight line) SSE of Ðuravci, near Besa / Bes near Krone i Besit , 330 m a.s.l., 42°08.548'N, 19°13.165'E (site code: 20171019G), leg. T. Deli, Z.P. Erőss, A. Hunyadi & B. Páll-Gergely, 19.10.2017, DT/60-70, EZP/60-70, HA/60-70, HNHM 103201/28, PGB/60-70 ; Montenegro, SE of Virpazar, 7.3 km S(SE) of Ðuravci, 1.6 km (in a straight line) NNW of Tejani , 455 m a.s.l., 42°06.803'N, 19°13.384'E (site code: 20171019I), leg. T. Deli, Z.P. Erőss, A. Hunyadi & B. Páll-Gergely, 19.10.2017, HNHM 103202/2 ; Montenegro, NNW of Virpazar, road between Rijeka Crnojevića and Virpazar, at the junction to Dupilo , 160 m a.s.l., 42°15.085'N, 19°04.987'E (site code: 20171020A), leg. T. Deli, Z.P. Erőss, A. Hunyadi & B. Páll-Gergely, 20.10.2017, HNHM 103203/1 ; Montenegro, S of Virpazar, 1.1 km (in a straight line) E of Limljani , one of the tunnels on the old road, 400 m a.s.l., 42°11.637'N, 19°06.309'E (site code: 20171021A), leg. T. Deli, Z.P. Erőss, A. Hunyadi & B. Páll-Gergely, 21.10.2017, DT/ca. 20, EZP/ca. 20, HA/ca. 20, HNHM 103204/6, PGB/ca. 20 ; Montenegro, S of Virpazar, 0.8 km (in a straight line) E of Limljani , above the village, 400 m a.s.l., 42°11.698'N, 19°06.217'E (site code: 20171021B), leg. T. Deli, Z.P. Erőss, A. Hunyadi & B. Páll-Gergely, 21.10.2017, DT/ca. 40, EZP/ca. 40, HA/ca. 40, HNHM 103205/8, PGB/ca. 40 ; Montenegro, NE of Bar, 1.2 km (in a straight line) NW of Tudjemili , 400 m a.s.l., 42°08.489'N, 19°08.144'E (site code: 20171021F), leg. T. Deli, Z.P. Erőss, A. Hunyadi & B. Páll-Gergely, 21.10.2017, DT/ca. 40, EZP/ca. 40, HA/ca. 40, HNHM 103206/10, PGB/ca. 40 ; Montenegro, 1.6 km (in a straight line) NE Petrovac, 0.3 km S of Novoselje , 470 m a.s.l., 42°13.032'N, 18°57.336'E (site code: 20171021G), leg. T. Deli, Z.P. Erőss, A. Hunyadi & B. Páll-Gergely, 21.10.2017, EZP/3 ; Montenegro, 2.8 km (in a straight line) NE Petrovac, 1.2 km E of Novoselje , 630 m a.s.l., 42°13.145'N, 18°58.245'E (site code: 20171021H), leg. T. Deli, Z.P. Erőss, A. Hunyadi & B. Páll-Gergely, 21.10.2017, DT/3, EZP/2, HA/4, HNHM 103207/1, PGB/2 ; Montenegro, Gradiste Monastery , 80 m a.s.l., 42°12.212'N, 18°57.772'E, leg. A. Reischütz, N. Reischütz & P. L. Reischütz, Jul. 2010, NHMW 112362/1 (photographed shell, Fig. 6F-J ); REI/5 ; Montenegro, above Sv. Stefan south of Budva , 110 m a.s.l., 42°14.926'N, 18°54.131'E, leg. A. Reischütz, N. Reischütz & P. L. Reischütz, Mar. 2011, REI/4 ; Montenegro, Tuđemili, 17 km towards Virpazar, Rumija Mts , 480 m a.s.l., 42°10.974'N, 19°6.546'E, leg. P. Subai, 15 Apr. 2009, NMBE 542070/56 ; Montenegro, Tuđemili, 1 km towards Virpazar, Rumija Mts , 395 m a.s.l., 42°8.238'N, 19°8.388'E, leg. P. Subai, 25 Sep. 2005, NMBE 542069/1 ; Montenegro, Tuđemili, 1 km towards Virpazar, Rumija Mts , 395 m a.s.l., 42°8.238'N, 19°8.388'E, leg. P. Subai, 25 Sep. 2005, NMBE 542068/3 ; Montenegro, Ðuravci, 10 Km E, southern side of Mount Kronistar , 500 m a.s.l., 42°6.594'N, 19°13.908'E, leg. P. Subai, 16 Sep. 2006, NMBE 542067/1 ; Montenegro, Zoganje N 5 km, on the Ulcinj-Shkodër road, 80 m a.s.l., 41°58.762'N, 19°15.547'E (site code: 2015/32), leg. T. Deli, Z.P. Erőss & Z. Fehér, 27 May 2016, DT/ca. 50, HNHM 103208/22, NHMW 112363/ca. 50 ; Montenegro, Rumija Mts, Virpazar S 9 km ( Virpazar-Bar road, between Boljevići and Tuđemili), old tunnel, 440 m, 42°11.474'N, 19°06.489'E (site code: 2008/182), leg. Z. Fehér, J. Kontschán & D. Murányi, 14.10.2008, HNHM 103209/23 ; Montenegro, scree (talus) ca. 5 km east of Dobra Voda in direction of Vladimir , 220 m a.s.l., 42°1.508'N, 19°11.160'E, leg. A. Reischütz, N. Reischütz & P. L. Reischütz, Jul. 2010, REI/6 ; Montenegro, Donji Murići, above the village, at the junction of a minor road to Besa , 200 m a.s.l., 42°9.233'N, 19°12.930'E (site code 2017/007), leg. Z.P. Erőss & Z. Fehér 16.07.2017, HNHM 103492/1 (juvenile shell, identification uncertain) .

Diagnosis.

A small to medium sized species with dense, low ribs, smooth protoconch, and toothless aperture.

Description.

Shell nearly flat, but spire somewhat always elevated; protoconch consists of 1.25-1.75 whorls, rather glossy; teleoconch with fine, equidistant, dense ribs; rib density variable (57-112 ribs on body whorl); between main ribs some fine wrinkles discernible; entire shell with 3.25-3.75 whorls; aperture semilunar; peristome slightly thickened and expanded; aperture toothless; umbilicus regular funnel-shaped, relatively narrow (width depends on spire height).

Measurements. SW: 1.9-3.4 mm (median = 2.3 mm), SH: 1.1-1.7 mm (median = 1.3 mm), AW: 0.7-1.3 mm (median = 0.9 mm), AH = 0.8-1.3 mm (median = 0.9 mm), AA = 56-68°(n = 15).

Differential diagnosis.

See under S. obodensis and S. hunyadii sp. n.

Variation among specimens.

This is a widely distributed species with numerous known populations, most of them with unique character states of spire height, shell size, and rib density.

Distribution.

This species is distributed in the Rumija Mountain between the Shkodër Lake Basin and the Adriatic Sea. Northwards the range extends to the Cetinje area. According to the original labels, type material is of Slovenian origin, however, this species was never again found in Slovenia. It can be reasonably supposed that it is due to mislabelling and the 'type locality’ is not the site where the type material actually came from ( Gittenberger 1975). Welter-Schultes (2012) reports the species only from Slovenia, which is based on the originally incorrect type locality.

Conservation status.

Assessed as Least Concern (LC) by Reischütz (2017a), because it is not an extremely rare species and there is no reason to suppose that the habitat quality, habitat extent or population are deteriorating or extremely fluctuating. Now, the number of known locations is more than 20, which confirms the LC status.

Figures

Figure 6. Shells of Spelaeodiscus dejongi Gittenberger, 1969. A-E holotype (NHMW 42517) F-J Montenegro, Gradiste Monastery (NHMW 112362) K-O Montenegro, 1.9 km (in a straight line) S of Virpazar (HNHM 103200).